Ancyropilus tricolor, Bartlett & Lambkin, 2022

Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, Zootaxa 5220 (1), pp. 1-81 : 21-24

Ancyropilus tricolor sp. nov.

ZooBank registration: urn:lsid:zoobank.org:act:4343EC1B-5A36-4041-8764-6A57456C00DD

( Figs 9 , 34 , 73 , 109 , 147 ; Map 1)

HOLOTYPE ♁: Western Australia: 22.30S 117.58E GPS, 28km NE of Tom Price , WA, 26 Sep 1995, D.C.F.Rentz & J.Otto, Stop 73 // 95-73 at light ( ANIC) . PARATYPES (28): Western Australia: WA, Pilbara region. Hamersley station, Horseshoe Bore , 300m N of windmill,—22:29:10, 117:41:28, 22-Sep-2005 to 27-Sep-2005, CVA Volunteers, PILB044/09P (pit trap) // 2023 // K 246875 (1, AM) ; WA, Pilbara region. Hamersley station, Horseshoe Bore , 300m N of windmill,—22:29:10, 117:41:28, 18-Nov-2004, 23-Nov-2004, LTM sites, CVA Volunteers, PILB044/03M [Malaise trap] // 1119 // K 246878 (1, AM); same data as previous except: 1117 // K 246866 (1, WAM E113533 , ex AM) ; WA, Pilbara region. Hamersley station, Horseshoe Bore , 300m N of windmill, -22:29:10, 117:41:28, 16- Feb-2005 to 20-Feb-2005, LTM sites, M. Bulbert & S. Ginn, PILB044/04M [] // 1116 // K 246867 (1, AM) ; same data as previous except: 1115 // K 246868 (1, WAM E113534 , ex AM) ; WA, Pilbara region. Hamersley station, Horseshoe Bore , 300m N of windmill, -22:29:10, 117:41:28, 13-Apr-2005 to 18-Apr-2005, LTM sites, M.Bulbert & G. Wood, PILB044/05M [Malaise trap] // 1112 // K 246871 (1 ♀, AM) ; same data as previous except: 1113 // K 246870 (1, WAM E113545 , ex AM) ; same data as previous except: 1114 // K 246869 (1 ♁, WAM E 113536 , ex AM) ; WA, Pilbara region. Hamersley station, Horseshoe Bore , 300m N of windmill, -22:29:10, 117:41:28, 30- Sep-2004 to 5-Oct-2004, LTM sites, CVA Volunteers, PILB044/02M [Malaise trap] // 207 // K 246876 (1 ♀, AM) ; WA, Pilbara region. Hamersley station, Horseshoe Bore , 300m N of windmill, -22:29:10, 117:41:28, 26-May-2004 to 29-May-2004, LTM sites, A. Donnelly & G. Carter, PILB044/01M [Malaise trap] // 2024 // K 246874 (1 ♀, WAM E113537 , ex AM) ; WA, Pilbara region. Juna Downs Station, Juna Downs Rd. to Packsaddle Bore approx. 20km E of homestead, -22:51:30, 118:40:14, 13-April-2005 to 18-April-2005, LTM sites, M. Bulbert & G. Wood, PILB036/05M [Malaise trap] // 1111 // K 246872 (1 ♁, WAM E113538 , ex AM) ; WA, Pilbara region. Juna Downs Station, Juna Downs Rd. to Packsaddle Bore approx. 5km E of homestead, -22:52:31 ; 118:31:49, 19-Nov-2004 to 24-Nov-2004, LTM sites, CVA Volunteers, PILB039/03M [Malaise trap] // 1118 // K 246879 (1, AM) ; WA, Pilbara region. Juna Downs Station, Great Northern Highway , approx. 8km S of Karijini Dr toff., -22:41:36, 118:42:19, 3-Jan-2006 to 8-Jan-2006, LTM sites, A. Donnelly & CVA Volunteer, PILB038/11M [Malaise trap] // 1751 // K 246865 (1, WAM E113539 , ex AM) ; WA, Pilbara region. Nanburra-Wittenoom Rd , approx 10km NE of Railway x-ing, nr railway crossing, -22:26:36, 117:48:23, 13-April-2005 to 18-April-2005, LTM sites, M. Bulbert & G. Wood, PILB042/05M [Malaise trap] // 1110 // K 246873 (1 ♁, AM) ; WA, 2km South-west of Paraburdoo along Paraburdoo-Nanutarra rd. , -23:11:53, 117:39:15, 19-Feb-2005, Adhoc collection, M. Bulbert & S. Ginn, PILB080/ LT6 // 1120 // K 246877 (1, AM); 22.25S 115.49E GPS, Timbuck Well , 35.5km ENE of Nanutarra H.S., WA, 7 Oct 1995, D.C.F.Rentz & L.Lowe, Stop 98 (1 ♀, ANIC) ; Marloo Stn. Wurarga, W.A., 1931-1941, A. Goerling (1, ANIC) ; Wurarga, WA, A. Goerling (1, ANIC) ; Australia,WA06/102, Dilgaty Brook, 22km nne. Landor-Carnarvon / Mullewa Rd 25.28591S, 116.21177E, 439m, 3.2.2006, M.Baehr (1, RGCM) ; Australia, WA06/118, Toolonga Cr. 30km ne. Cane River , 21.95651S, 115.65010E, 89m, 10.2.2006, M.Baehr (1, WAM E113541 , ex RGCM) ; 11 mi. SbyW of Cocklebiddy, WA, 32.45S 126.03E, 22.xi.69, at light, Mallee, Callitris, on white sand, Britton, Taylor & Upton (1, ANIC) ; 26.03S 127.14E, 66km EbyN of Warburton , WA, 15.xi.1977, T.A.Weir (1 ♁, 1 ♀, 1, ANIC) . South Australia: 27.18S 133.41E, 41 km WbyN of Welbourn Hill , SA, 21.xi.1977, T.A. Weir (1 ♀, ANIC) ; S.Aust., Vokes Hill Junc., 30°8′S 135°4′E, at light, Sandy Desert surv, 19 Sept 2001, PJ Lang (1 ♁, SAMA) ; S.Aust., Gawler Ra., Kolay Dam, 21 33′S 135 36′E, at light, Paney Stn. , 7 Dec 1989, J. Forrest (1, SAMA) . Queensland: Qld: 28.872° Sx 144.502°E, Currawinya NP, Woolshed (CW1 M), 1-12 Dec 2008, 136m, Malaise, Mulga A. Townsend J. Burke, 18116 (1 ♁, QM) .

Diagnosis. Pronotum rounded, disc densely punctate, with three small glabrous regions (one baso-medially, one either side of central impression); elytra dark with brown and yellowish maculations (each elytron with a brown basal region, a small yellowish baso-lateral macula beneath humeral angle, a yellow transverse fascia curved before suture and an angular apical macula), punctation without nodules, 8 th stria beginning near base (between 2 nd and 7 th punctation of 7 th stria), all striae reaching apical macula, occasionally more than one seta across interstrial width; femora yellow and brown (femora of some specimens with black also), tarsi with three ventral tarsal pads.

Ancyropilus tricolor sp. nov. differs from the morphologicaly similar A. brigalowae sp. nov. by the V-shaped internal margin of the tegminal dorsal sinus and the colouration of the pronotum and elytra base (brown rather than black).

Description. Habitus: Fig. 147 . Total length: 4.4–7 mm (holotype, 6.8 mm). Head: Cranium dark brown, clypeus and supra-antennal elevations more reddish, anteclypeus semi-transparent orange, labrum, antennae and palpi orange to orange-brown; eyes separated by about 0.6–0.87 eye widths (holotype, 0.77); vertex heavily rugulose, upper part of frons more punctate than rugulose, lower part of frons (above epistomal suture) less punctate with light transverse rugulosity, slightly raised on some specimens; clypeus not heavily punctate; genae and submentum wrinkled; exterior margins of terminal palpomeres about 1.4 times (maxillae) and 2 times (labium) the length of inside edges; antennae almost reaching base of pronotum; eyes and cranium with well-separated long erect pale setae, frons and vertex with denser shorter medially-directed setae. Prothorax: Brown (castaneous); pronotum 1.13–1.18 times longer than wide (holotype, 1.14), sides rounded, middle slightly wider than pronotal arch; subapical depression deeply v-shaped; central impression short, punctate; disc convex, dense with irregularly-shaped punctation, a longitudinal glabrous stripe basally, an ovoid glabrous tumescence either side of central impression; well distributed with fine short setae and sparsely with longer erect setae. Pterothorax: Ventrites brown, with fine pale posteriorly- or medially-directed setae; elytra blackish-brown with yellowish and brown markings (each elytron with a yellowish apical angulate macula, a broad yellowish central fascia curved at the suture, a small yellowish baso-lateral macula beneath humeral angle and a brown scalene-triangular basal region—the two longest sides running from the humerus to the suture and along the suture), length to width ratio 2.47–2.74:1 (holotype, 2.56:1); 8 th stria beginning anterior of fascia, between 2 nd and 7 th punctation of 7 th stria (first few punctations widely spaced in some specimens), all striae extending to apical macula; punctation circular, without nodules; epipleurae extending into apical maculae; interstriae with short fine setae (at least one per puncture) along sides of striae (proximal to edge of punctations), plus longer thicker erect setae (at least one every two punctures) along midline of striae (more than one seta across strial-width when the two setal types overlap); hindwing with CuA 3+4 and CuA 1 cross-veins complete, MP 3+4 basad CuA 1 crossvein small (but not completely absent). Legs: Approximately basal one-third to two-fifths of profemora and basal half of meso- and metafemora yellow, remaining parts of femora black-brown centrally and brown apically (or just brown in pale specimens); tibiae and tarsi brown, tibial carinae darker; ventral tarsal pads yellowish; profemora moderately swollen, other femora slender. Abdomen: Ventrites light to dark brown (tan). Male genitalia: Tegminal plate ( Figs 9 , 34 ) with base slightly broader than parameroid lobes, weakly narrowing between middle and parameroid lobes or not, apex externally tapered toward sinus, not acuminate or digitiform, dorsal sinus about one-fifth tegmen length, internal limit more angular than rounded, inner margins interrupted by opening above middle, ventral sinus slightly longer, tegminal arms tapering gradually to meet broad apodeme, length of apodeme about one-fifth to one-quarter tegmen length (difficult to determine); median lobe as in Fig. 73 ; pygidium as in Fig. 109 .

Etymology. The specific epithet tricolor refers to the three-tone pigmentation of the elytra (black, brown and yellow).

Biology. Specimens of the type series were collected from September to May at light or using Malaise or pit traps.

Distribution (Map 1). Ancyropilus tricolor sp. nov. is described from specimens collected in Western Australia, South Australia, and southern Queensland.

Figures

FIGURES 1–11. (1) Opilo mollis, elytron (1a) first interval broad, (1b) pre-apical ridge; (2–9) tegmina, lateral—(2) Opilo mollis (arrow showing weakly sclerotised tegminal arm region); (3) Notopilo cf. lawnhillensis; (4) Notopilo gerstmeieri, paratype; (5) Notopilo obesus, paratype; (6) Infectostriatus differens, holotype; (7) Notopilo congruus; (8) Culcipternulus mareebensis, paratype; (9) Ancyropilus tricolor, paratype; (10–11) antennal scape—(10) Opilo mollis (carina absent); (11) Notopilo congruus (arrow showing carina) (scale bars: Figs 1–9, 0.5 mm; Figs 10–11, 0.1 mm).

FIGURES 27–47. Tegmina, dorsal: (27) Ancyropilus brigalowae, paratype; (28) Ancyropilus emmotti, holotype; (29) Ancyropilus exossuarius, paratype; (30) Ancyropilus monteithi, holotype; (31) Ancyropilus noonbahensis, holotype; (32) Ancyropilus packsaddlensis, holotype; (33) Ancyropilus simplex, paratype; (34) Ancyropilus tricolor, paratype; (35) Ancyropilus hornensis, paratype; (36) Culcipternotus mareebensis, paratype; (37) Infectostriatus absentis, holotype; (38) Infectostriatus differens, holotype; (39) Monilonotum bunyense, paratype; (40) Monilonotum doddi; (41) Monilonotum pascoei; (42) Monilonotum sundholmi, paratype; (43–47) Notopilo, beswickensis species group—(43) Notopilo beswickensis, holotype; (44) Notopilo calicis, paratype; (45) Notopilo tompricensis, holotype; (46) Notopilo xanthoimprocerus, holotype; (47) Notopilo xanthoprolatus, holotype (scale bars: 0.5 mm).

FIGURES 66–101. Median lobes: (66) Ancyropilus brigalowae, paratype; (67) Ancyropilus emmotti, holotype; (68) Ancyropilus exossuarius, paratype; (69) Ancyropilus monteithi, holotype; (70) Ancyropilus noonbahensis, holotype; (71) Ancyropilus packsaddlensis, holotype; (72) Ancyropilus simplex, paratype; (73) Ancyropilus tricolor, holotype; (74) Ancyropilus hornensis, paratype; (75) Culcipternotus mareebensis, paratype; (76) Infectostriatus absentis, holotype; (77) Infectostriatus differens, holotype; (78) Monilonotum bunyense, paratype; (79) Monilonotum pascoei; (80) Monilonotum sundholmi, paratype; (81–85) Notopilo, beswickensis species group—(81) Notopilo beswickensis, holotype; (82) Notopilo calicis, paratype; (83) Notopilo tompricensis, holotype; (84) Notopilo xanthoimprocerus, paratype; (85) Notopilo xanthoprolatus, holotype; (86–88) Notopilo, cambageicola species group—(86) Notopilo cambageicola, paratype; (87) Notopilo interfabulatus, paratype; (88) Notopilo tanybasilaris, paratype; (89) Notopilo congruus (congruus species group); (90) Notopilo reduncus (reduncus species group); (91–93) Notopilo, variipes species group—(91) Notopilo brevistriatus, paratype; (92) Notopilo gerstmeieri, holotype; (93) Notopilo variipes; (94–96) Notopilo, unplaced to species group—(94) Notopilo elstoni, holotype; (95) Notopilo katherinensis, holotype; (96) Notopilo obesus, paratype; (97) Platynotum bulli, paratype; (98) Platynotum culgoense, holotype; (99) Platynotum femorale; (100) Platynotum foveosetosa, holotype; (101) Platynotum gracile, holotype (scale bars: 0.5 mm).

FIGURES 102–137. Male pygidiae: (102) Ancyropilus brigalowae, paratype; (103) Ancyropilus emmotti, holotype; (104) Ancyropilus exossuarius, paratype; (105) Ancyropilus monteithi, holotype; (106) Ancyropilus noonbahensis, holotype; (107) Ancyropilus packsaddlensis, holotype; (108) Ancyropilus simplex, paratype; (109) Ancyropilus tricolor, paratype; (110) Ancyropilus hornensis, paratype; (111) Culcipternotus mareebensis, paratype; (112) Infectostriatus absentis, holotype; (113) Monilonotum doddi; (114) Monilonotum pascoei; (115) Monilonotum sundholmi, paratype; (116–120) Notopilo, beswickensis species group—(116) Notopilo beswickensis, holotype; (117) Notopilo calicis, paratype; (118) Notopilo tompricensis, holotype; (119) Notopilo xanthoimprocerus, paratype; (120) Notopilo xanthoprolatus, holotype; (121–124) Notopilo, cambageicola species group—(121) Notopilo cambageicola, paratype; (122) Notopilo interfabulatus, paratype; (123) Notopilo lawnhillensis, paratype; (124) Notopilo tanybasilaris, holotype; (125) Notopilo congruus (congruus species group); (126–128) Notopilo, variipes species group—(126) Notopilo brevistriatus, paratype; (127) Notopilo gerstmeieri, holotype; (128) Notopilo variipes; (129–132) Notopilo, unplaced to species group—(129) Notopilo confusus, holotype; (130) Notopilo elstoni, holotype; (131) Notopilo katherinensis, holotype; (132) Notopilo obesus, paratype; (133) Platynotum bulli, paratype; (134) Platynotum culgoense, holotype; (135) Platynotum femorale; (136) Platynotum foveosetosa, holotype; (137) Platynotum gracile, holotype (scale bars: 0.5 mm).

FIGURES 138–149. Habitus: (138) Ancyropilus brigalowae, paratype; (139) Ancyropilus emmotti, holotype; (140) Ancyropilus exossuarius, paratype; (141) Ancyropilus hornensis, paratype; (142) Ancyropilus labris, holotype; (143) Ancyropilus monteithi, paratype; (144) Ancyropilus noonbahensis, holotype; (145) Ancyropilus packsaddlensis, holotype; (146) Ancyropilus simplex, paratype; (147) Ancyropilus tricolor, paratype; (148) Culcipternotus mareebensis, holotype; (149) Infectostriatus absentis, holotype (scale bars: 1.0 mm).