Umenocoleidae Chen et Tan, 1973

Sendi, Hemen, 2024, Nigropterix cummingi sp. n., an umenocoleid (Insecta: Dictyoptera) from mid-Cretaceous northern Myanmar amber, Zootaxa 5424 (4), pp. 476-482 : 477

Family Umenocoleidae Chen et Tan, 1973

Type species. Umenocoleus sinuatus Chen et T’an, 1973

Stratigraphic range. Latest Aptian-Turonian.

Geographic range. Cosmopolitan, both Laurasia and Gondwana.

Composition. Umenocoleus Chen et Tan, 1973 ; Ponopterix Vršanský et Grimaldi, 1999 ; Blattapterix Vršanský, 2003 ; Elytropterix Vršanský, 2003 ; Petropterix Vršanský, 2003 ; Compunctiotypus Kaddumi, 2005 ; Umenopterix Lee, 2016 ; Enervipraeala Luo, Xu et Jarzembowski, 2021 ; Laticephalana Luo, Beutel, Xu et Jarzembowski, 2021 ; Archaeospinapteryx Sendi et Cumming, 2023; Trapezionotum Sendi et Cumming, 2023; Poikiloprosopon Sendi et Cumming, 2023; Nigropterix Sendi, 2023 ; Umenotypus Sendi, 2023. Alienopterix Mlynský, Vršanský et Wang, 2018 ; Nadveruzenie Vršanský, Hinkelman et Sendi, 2021 ; Vzrkadlenie Vršanský, 2021 ; Jantaropterix Vršanský et Grimaldi, 2003 ; Perspicuus Koubová, 2020 ; Lepidopterix Sendi, 2020 ; Antophiloblatta Sendi, 2020 ; are transferred to Umenocoleidae (see also Sendi et al. 2023).

Diagnosis (after Luo et al. 2022; Vršanský 2003; Nel et al. 2014; Lee 2016; Sendi et al. 2023). Head transverse, orthognathous. Compound eyes large, oval. Antenna sometimes thin with numerous wide segments and long setae in transverse rows. Pronotum sometimes subdivided by a transverse supracoxal furrow. Forewings sclerotized, often covered by cup-shaped punctures (‘‘bunky”). Venation simplified or sometimes indistinct, adsutural line sometimes present. Distal part of hindwings distinctly projected beyond forewing apex. Stem R reduced, with few branches, possibly with pterostigma in R1. Stem M branched, stem CuA with many branches and stem CuP simple. Cerci sometimes with long setae and few segments. Females with short external ovipositor.