Rhamphomyia (Holoclera) helvetica, Barták & Kubík, 2012

Barták, Miroslav & Kubík, Stepán, 2012, A review of the Palaearctic species of Rhamphomyia subgenus Holoclera (Diptera: Empididae) with description of 5 new species, Revue suisse de Zoologie 119 (3), pp. 385-407 : 392-395

Rhamphomyia (Holoclera) helvetica sp. n. Fig. 3

HOLOTYPE: MHNG; SWITZERLAND, GE, Cartigny / Moulin de Vert; 1 Ƌ; 350 m, 5.vi.2006, B. Merz.

PARATYPES: MHNG, CULSP; SWITZERLAND. 5 Ƌ, 2 ♀; same data as the holotype .- 1 ♀; same locality; 2.vi.2002.- 1 Ƌ; same locality, 470 m; 4.vi.2001; M. Eggenberger & B. Merz. - 1 Ƌ; Chancy / LaLaire , 350 m; 1.vii.2001; B. Merz. - 1 Ƌ , 2 ♀; Cartigny , Nant-des-Crues , Rhone embouchure, 360 m; 4.vi.2001; M. Eggenberger & B. Merz. - 1 Ƌ; Dardagny / LeMoulin, 360 m; 30.vi.2001; B. Merz .- 1 ♀; Dardagny / Roulave, 420 m; 30.vi.2001; B. Merz .- 1 ♀; Avusy Moulin de la Grave , MT; 7.vi.2006; H. Boillat .- 1 ♀; Russin , Les Baillets, 405 m; 30.vi.2001; B. Merz. - 3 Ƌ; NE, Marin / Les Tertres, 435 m; 19.v.2001; B. Merz .- 1 ♀; St. Blaise / Les Riedes, 470 m; 19.v.2001; B. Merz. - 1 Ƌ; ZH, Dietikon-Hardwald, 400 m; 15.v.1995; B. Merz. - 1 Ƌ; Zürich-Allmend, 450 m; 8.vi.1994; B. Merz .- UZMC; SPAIN. 1 Ƌ; Granada , Rio Mulhacén, 5 km N Capileira, 1500 m; 9. iv 1966 .- 1 ♀; Rio Guadalfeo , Orgiva, 300 m; 4. iv 1966; Lyneb. - Martin-Langem .

Rhamphomyia (Holoclera) helvetica sp. n., male terminalia (macerated), lateral view. Scale 0.1 mm.

DISTRIBUTION: Spain, Switzerland.

DATE OF OCCURRENCE: April-July.

DESCRIPTION

Male: Eyes holoptic, facets in ventral half of eye much smaller than in dorsal half. Frons (small area just above antenna and below ocellar triangle) brown, brown microtrichose, setae absent. Ocellar setae black, about half as long as the distance between front ocellus and base of antenna, ocellar triangle with 1 pair of additional setae on hind margin. Face brown, microtrichose, about 0.09-0.10 mm broad ventrally and 0.11-0.14 mm long, without setae. Occiput dark brown, microtrichose, sparsely black setose (setae on dorsal half of occiput arranged in two rows), postocular row irregular on ventral half. Antenna brown, length of antennal segments (in 0.01 mm scale) = 5-6: 7: 21-26: 10-12, both basal antennomeres short setose (the longest setae about 0.07 mm long). Labrum brown, slightly shorter than head is high. Palpus brown, short, with 2-3 short setae. Gena very narrow, clypeus microtrichose. Thorax brownish black, microtrichose, mesoscutum uniformly dark brownish black, almost velvety black in dorsal view; without any stripes. All thoracic setae black. Chaetotaxy: 0-3 setae on proepisternum; both prosternum and proepisternal depression bare; 7-10 uniserial dorsocentrals (about 0.15 mm long in middle of their rows), 1-2 only slightly longer inclinate prescutellars; acrostichals absent; intrahumeral absent; one strong posthumeral (no additional setae laterad of dorsocentrals in presutural part of mesoscutum); 1 long postpronotal and 1-2 short additional setae; 2-3 notopleurals (0-2 shorter setae on anterior part of notopleuron); both supraalars and prealars absent; 1 strong and 1 very small postalars; 4 scutellars; laterotergite with black setae. Legs including coxae brown (fore coxa yellowish brown apically, yellow in the male from Spain), microtrichose, hind femur (in some specimens also tibiae) lustrous, all setae black. No seta in posteroapical comb on hind tibia. Fore femur with short setation (no prominent setae), anterior row of (sensory?) setae 0.02-0.04 mm long. Fore tibia with short setation only, without prominent setae (except preapicals). Mid femur with short setation, no prominent setae except preapicals. Mid tibia often with one anteroventral seta on apical third, otherwise with short setation only. Hind femur with short setation, without prominent setae, except several preapical anteroventrals not much stronger than other setation but somewhat longer. Hind tibia slightly broadened apically, with 1-3 posterodorsal setae slightly longer than tibia is deep (situated mostly on distal half of tibia), otherwise with short setation only. Basal tarsomeres of all legs thin and short setose, ventral setae short, more distinct on hind one. Measurements (in 0.01 mm scale): length: f1 (72-85), f2 (83-95), f3 (115-140), t1 (75-88), t2 (79-88), t3 (105-120), bt1 (41-50), bt2 (35-38), bt3 (48-58), width: bt1 and bt2 (5), bt3 (6-7). Wing light brown, stigma indistinct or only slightly darkened, veins brown, anal vein (A1) indistinct in distal third. Costal seta long, axillary angle right. M2/D = 1.8-2.3, CuA1 ratio = 3.4-5.8, lw: ww = 2.9-3.3. Halter dark brown, calypter brown with dark fringes. Abdomen brown (lighter ventrally and almost velvety blackish brown in dorsal view), brown microtrichose, black setose. Hind marginal setae on sides of tergites 2-4 subequally long as their corresponding segments, on remaining segments slightly shorter, discal setae shorter than marginals, dorsum of tergites with short setae, 1st sternite bare. Terminalia ( Fig. 3 ): hypandrium triangle-shaped in lateral view, with several setae, basal part connected with phallobase with very long and narrow arms; cercus short and simple, straigth dorsally; epandrium rounded apically, with a single spine at apex; phallus medium long and evenly thin to the tip. Length of body 2.5-3.1 mm, wing 2.3-3.2 mm.

Female: Similar to male but with the following differences. Eyes dichoptic, facets subequal in size. Frons brown, microtrichose, 0.18-0.25 mm long and 0.08-0.09 mm broad, almost parallel sided, with 3-6 rather long (up to 0.10 mm) setae on each side. Face subequally broad as frons. Ocellar setae ¾ as long as the distance between front ocellus and base of antenna. Mesoscutum dark reddish brown to blackish brown, sublustrous. Legs paler than in male: fore coxa yellow, mid and hind coxae yellow to brownish yellow, proximal parts of all femora yellow to brownish yellow. Basal tar- someres of legs with slightly longer ventral spines than in male. Measurements (in 0.01 mm scale): length: f1 (72-95), f2 (80-100), f3 (110-125), t1 (70-88), t2 (78-94), t3 (105-117), bt1 (38-45), bt2 (32-36), bt3 (44-52), width: bt1 and bt2 (4), bt3 (5-6). Wing light brownish, M2/D = 1.6-1.9, CuA1 ratio = 2.6-4.8, lw: ww = 2.7-2.8. Halter brown, calypter brown with dark fringes. Abdomen yellow to brownish yellow proximally and dark blackish brown distally, microtrichose, lateral part of tergites 2-4 sublustrous. Hind marginal setae on sides of tergites 2-4 less than half as long as their corresponding segments, on remaining segments slightly shorter, discal setae slightly shorter than marginals. Length of body 2. 0-3.1 mm, wing 2.6-3.4 mm.

DIFFERENTIAL DIAGNOSIS: R. (Holoclera) helvetica sp. n. belongs to the R. (Holoclera) nigripennis group and it is most similar to R.­ nigripennis (Fabricius) (both species sharing two characteristics: supraalar seta absent and cercus almost straight dorsally). The R.­ nigripennis male has a very long phallus (see Barták 1982: fig. 12d), much longer than in R.­ helvetica . However, this characteristic is visible only after dissection. The shape of cercus may be used for distinguishing both species without dissection: in R.­ nigripennis it has distinct basal swelling, which is best visible in dorsal view (inner margins are angled) but it has no swelling in R.­ helvetica (inner margins of cerci are straight, V-shaped in dorsal view). Moreover, epandrium is narrowed apically in R.­ helvetica with only a single apical spine, but broadened apically with several long spine-like setae in R.­ nigripennis . Female frons is microtrichose in R.­ helvetica but lustrous in R.­ nigripennis .

DERIVATIO NOMINIS: the specific name is derived from the country of origin

(Helvetia = Switzerland).

Figures

FIG. 3