Ischnothyreus ligulatus Richard,

Richard, Miguel, Graber, Werner & Kropf, Christian, 2016, The goblin spider genus Ischnothyreus (Araneae, Oonopidae) in Java and Sumatra, Zootaxa 4151 (1), pp. 1-99: 17-26

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Ischnothyreus ligulatus Richard

sp. nov.

Ischnothyreus ligulatus Richard  , sp. nov.

( Figs. 9–16View FIGURE 9View FIGURE 10View FIGURE 11View FIGURE 12View FIGURE 13View FIGURE 14View FIGURE 15View FIGURE 16)

Type material. Holotype: male (PBI_OON00052392), JAVA: Ujung Kulon National Park (6° 44′ 48″ S, 105° 20′ 1″ E) November 14–25, 1986, Leg. Suh. Djojosudharmo et al., coll. Christa Deeleman, NNM Leiden, NLGoogleMaps  . Paratypes: 6 males and 8 females (PBI_OON00031890), collected with holotype, NNM Leiden.

Additional material examined. 5 males and 6 females, collected with holotype, NNM Leiden. 1 female (PBI_OON00012053), JAVA: W Java, Mt Gede , 50km SE Bogor, 2600m, November 5, 1989, Agosti, Löbl, Burckhardt # 5a, MHNG  . 2 males, 6 females (PBI_OON00031897), JAVA: W  . Java, Cibodas , 1500–1600m, December 1986, leg. Suh. Djojosudharmo, Coll. Deeleman, NNM Leiden  . 6 males, 2 females (PBI_OON00031886), JAVA: Ujung Kulon National Park (6° 44′ 48″ S, 105° 20′ 1″ E), November 14–25, 1986, leg. SuhGoogleMaps  . Djojosudharmo , Coll. Deeleman, NNM Leiden  . 1 female (PBI_OON00031899), JAVA: Ujung Kulon National Park , leaflitter, November 21, 1986, leg. Suh  . Djojosudharmo , coll. Christa Deeleman, NNM Leiden. 

Etymology. The specific name is an adjective and refers to the spoon-like structures on the base of the male cheliceral fang and the tip of the embolus. It is derived from the latin word for spoon, ligula.

Diagnosis. Males of I. ligulatus  sp. nov. resemble I. florifer Kranz-Baltensperger, 2011  males as they have very similar pedipalps with a spoon-shaped, less sclerotized appendage on the embolic tip (ssa, Figs. 10View FIGURE 10 E, F, 12E, F). They can be seperated by the absence of the spoon-shaped process on the cheliceral fang in I. florifer  . The bulb has a single large ventral protuberance ( Figs. 10View FIGURE 10 G, H).

Females of I. ligulatus  sp. nov. resemble I. barratus Edwards & Harvey, 2014  and I. monteithi Edwards & Harvey, 2014  with small lateral sclerites flanking the postepigastric scutum (ls, Fig. 15View FIGURE 15 A) and a roundish appearance of the winding tube. But the flanking apodemes are different and the nail-shaped process on the postepigastric scutum (wt, a, nsp, Figs. 15View FIGURE 15 A –E, 16E –G) is lacking in the other species.

Description. Male: (PBI_OON00031890): Total length 1.65; carapace length: 0.79; carapace width: 0.65; abdomen length; 0.86.

CEPHALOTHORAX: Carapace pale orange, ovoid in dorsal view, pars cephalica strongly elevated in lateral view, anteriorly narrowed to between 0.5 and 0.75 times its maximum width, surface of elevated portion of pars cephalica smooth, sides smooth; lateral margin straight, smooth, present in one row. Clypeus margin unmodified, straight in front view, vertical in lateral view, high, ALE separated from edge of carapace by their radius or more; setae dark, needle-like. Eyes six, well developed, ALE largest, ALE circular, PME squared, PLE oval; posterior eye row straight from above, procurved from front; ALE touching, ALE-PLE separated by less than ALE radius, PME touching throughout most of their length, PLE-PME touching. Sternum as long as wide, pale orange, uniform, surface smooth, microsculpture absent, lateral margin without infra-coxal grooves, distance between coxae approximately equal, extensions of pre-coxal triangles present, lateral margins unmodified; setae dark, needle-like, densest laterally, originating from surface. Mouthparts: Chelicerae, endites and labium pale orange.

Chelicerae straight, anterior face unmodified; fangs directed posteriorly, shape normal, fang base with spoonshaped process (p, Figs. 11View FIGURE 11 C, D, 13B –D); tip unmodified; setae light, needle-like, evenly scattered; paturon inner margin with scattered setae. Labium rectangular, not fused to sternum, anterior margin not indented at middle, same as sternum in sclerotization; with 3–5 setae on anterior margin, subdistal portion with unmodified setae. Endites distally not excavated, serrula absent, anteromedian tip with one strong, tooth-like projection, posteromedian part unmodified, same as sternum in sclerotization, with anterolateral, strongly sclerotized toothlike projection.

ABDOMEN: cylindrical, rounded posteriorly; dorsum soft portions white, without color pattern. Posterior spiracles not connected by groove. Scutum not extending far dorsal of pedicel, cuticular outgrowths near pedicel absent. Dorsal scutum weakly sclerotized, yellow-brown, without color pattern, covering more than 3/4 of abdomen, more than 1/2 to most of abdomen width, not fused to epigastric scutum. Epigastric scutum weakly sclerotized, smooth, surrounding pedicel, not protruding. Postepigastric scutum weakly sclerotized, pale orange, long, almost rectangular, covering about 1/2 of abdominal length, not fused to epigastric scutum. Dorsum setae dark, needle-like. Epigastric area setae uniform, dark, needle-like. Postepigastric area setae dark, needle-like. Spinneret scutum with fringe of needle-like setae. Interscutal membrane with setae.

LEGS: yellow, without color pattern; femur IV same size as femora I-III, patella plus tibia I near as long as carapace, tibia I unmodified. Leg spination: femora: I p0-1-1; II p0-0-1; tibiae: I, II p2-1-1; r2-1-1; metatarsi: I, II p1-1-0; r1-1-0.

GENITALIA: Epigastric region with sperm pore large, oval, situated at level of anterior spiracles, unmodified; without setae. Bulb with one large ventral protuberance, retrolateral lobe simple. Tip of bulb slightly elongated, spoon-like, with spoon-shaped less-sclerotized appendage (ssa, Figs. 10View FIGURE 10 E, F, 12E, F).

Female: (PBI_OON00031890). Total length 1.71 carapace length: 0.79; carapace width: 0.70; abdomen length; 0.98.

As in male except as noted.

CEPHALOTHORAX: Female palp claws absent; spines formula: femur p1-1-1. Serrula present. Endites unmodified.

ABDOMEN: ovoid, pointed posteriorly; dorsum soft portions yellow-brown. Dorsal scutum orange-brown, covering about 1/2 of abdomen. Epigastric scutum small, lateral sclerites present. Postepigastric scutum yellowbrown, widely hexagonal, only around epigastric furrow, with short posteriorly directed lateral apodemes, characteristic posteriorly pointing nail-shaped projection on ventral surface; may be hardly visible because situated beneath darkly sclerotized winding tube. LEGS: pale orange; patella plus tibia I longer than carapace. Leg spination: femora: I p0-1-1; II p0-0-1; tibiae: I p2-1-1; r2-1-1; II p2-1-1; r2-2-0; metatarsi: I, II p1-1-0; r1-1-0. Proclaws inner face smooth.

GENITALIA: strongly sclerotized winding tube with globular appearance, apodemes lateral of winding tube with openings to surface of postepigastric scutum. Postepigastric scutum with central posteriorly pointing nailshaped process (wt, a, nsp, Figs. 15View FIGURE 15 A –E, 16E –G). Row of small pores from anterior middle of postepigastric scutum to base of nail-shaped process (po, Fig. 16View FIGURE 16 F).

Distribution. Only known from Ujung Kulon National Park in western Java.


National Museum of Natural History, Naturalis


Museum d'Histoire Naturelle