Landouria tonywhitteni, Nurinsiyah & Neiber & Hausdorf, 2019

Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, European Journal of Taxonomy 526, pp. 1-73 : 45-47

publication ID	https://doi.org/ 10.5852/ejt.2019.526
publication LSID	lsid:zoobank.org:pub:3FFC527D-E9CA-4D9F-BF59-076FA60171AC
persistent identifier	https://treatment.plazi.org/id/59260748-3438-411F-82B4-14A7847AFA58
taxon LSID	lsid:zoobank.org:act:59260748-3438-411F-82B4-14A7847AFA58
treatment provided by	Plazi
scientific name	Landouria tonywhitteni
status	sp. nov.

Treatment

Landouria tonywhitteni View in CoL sp. nov.

urn:lsid:zoobank.org:act:59260748-3438-411F-82B4-14A7847AFA58

Figs 39 View Figs 29–47 , 52 View Figs 52–55 , 60 View Figs 60–62 , 63 View Figs 63–71 ; Tables 3–4 View Table 3 View Table 4

Diagnosis

Landouria tonywhitteni sp. nov. is characterized by having a strongly depressed conical, brownishcorneous, sharply keeled shell and a penis that does not become distinctly broader towards is proximal end, the lack of a coecum at the proximal end of the penis and the presence of a node at the base of the flagellum.

Etymology

Landouria tonywhitteni sp. nov. is named in honour of the late British naturalist Tony Whitten (1953– 2017), who dedicated his life to the conservation of karst habitats and especially the snails that inhabit these habitats.

Material examined

Holotype

INDONESIA • holotype (det. anat.); Central Java, Grobogan, Taruman, near Sendang Sedayu ; 7°00′52″ S, 110°51′29″ E; 20 m a.s.l.; 16 Aug. 2014; A.S. Nurinsiyah, F.L.H. Irsyad, F.J. Sari and E. Nurlaela leg.; measurements: D = 17.6 mm, H = 7.6 mm; MZB 19214. GoogleMaps

Paratypes

INDONESIA – Central Java • 9 spec.; same data as for holotype; MZB 19217 GoogleMaps • 5 spec.; same data as for holotype; ZMH 133434 View Materials GoogleMaps • 9 spec. (det. anat.); Grobogan, Taruman, near Sendang Sedayu; 7°00′52″ S, 110°51′29″ E; 23 m a.s.l.; MZB 19215 • 1 spec.; same data as for preceding; MZB 21268 GoogleMaps • 5 spec.; same data as for preceding; ZMH 133436 View Materials GoogleMaps .

Description

SHELL ( Fig. 60 View Figs 60–62 ; Tables 3–4 View Table 3 View Table 4 ). Depressed conical, with 5.75–6 hardly convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles; apical side with indistinct spiral lines, umbilical side with indistinct incised spiral lines; with radial scaly processes; tuberculate around umbilicus; brownishcorneous; body whorl sharply keeled; aperture rhombic; upper insertion of peristome slightly descending; peristome expanded, reflexed and slightly thickened; umbilicus slightly eccentric, wide, comprising 32–41% of shell diameter, hardly obscured by columellar edge.

GENITALIA ( Figs 39 View Figs 29–47 , 63 View Figs 63–71 ; Table 4 View Table 4 ). Atrium short; penis long, consisting of a long, cylindrical distal part covered by an additional sheath, which is connected by muscular tissue to proximal part of epiphallus and a dilated proximal part containing a short penial papilla; without penis coecum; penial retractor runs from diaphragm to distal part of long, narrow epiphallus; flagellum short, broadest at its base, where it is equipped with a globular node, tapering towards its proximal end, not distinctly crenated; vas deferens narrow and long; vagina long; peduncle of bursa copulatrix subdivided into a slightly broader, cylindrical distal part and a narrower, longer proximal part, ending in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina.

Remarks

Landouria nodifera sp. nov., L. tonywhitteni sp. nov., L. pacitanensis sp. nov., L. sewuensis sp. nov. and L. intumescens share strongly depressed, brownish-corneous, sharply keeled shells. These species cannot be reliably distinguished by shell characters. Landouria tonywhitteni sp. nov. shares a node at the base of the flagellum with L. nodifera sp. nov., from which it differs in the lack of a coecum at the proximal end of the penis and in having a penis that does not become broader towards its proximal end.

Distribution

Landouria tonywhitteni sp. nov. is endemic to Sukolilo Karst (Fig. 52), where it occurs sympatrically with L. sukoliloensis sp. nov.

Figures

Figs 29–47. Flagella of species of Landouria from Java. 29. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 30. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 31. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 32. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 33. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 34. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 35. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 36. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 37. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 38. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). 39. L. tonywhitteni sp. nov., Central Java, Taruman Village, near Sendang Sedayu (holotype, MZB 19214). 40. L. nodifera sp. nov., Central Java, Larangan Village, Lanang Spring (holotype, MZB 19227). 41. L. pacitanensis sp. nov., East Java, Pacitan, near Srau Beach (MZB 19231). 42. L. intumescens (Martens, 1867), East Java, Guwoterus (MZB 19207). 43. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). 44. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 45. L. ciliocincta (Möllendorff, 1897), East Java, Malang, Kondangmerak (MZB 19211). 46. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 47. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bar: 1 mm.