Mycomya (Neomycomya) Väisänen

Amorim, Dalton De Souza & Oliveira, Sarah Siqueira, 2013, Types of Neotropical Mycetophilidae (Diptera) at the Natural History Museum collection, London, Zootaxa 3726 (1), pp. 1-119 : 57-62

Mycomya (Neomycomya) Väisänen

Mycomya (Neomycomya) Väisänen 1984: 292 . Type-species: Sciophila fimbriata Meigen 1818 (orig. des.).

M. alexanderi ( Coher) 1950b: 565 , fig. 2 (♂ terminalia). Type-locality: Brazil, São Paulo, Salesópolis, Estação Biológica de Boracéia. Holotype ♂, MZUSP.

Notes. The type series included the male holotype and fourty paratypes without gender especification, of which eight male paratypes, all from the type-locality, are at the NHM ( BMNH (E) # 256648-54 / 256656) .

M. bequaerti ( Coher) 1950b: 577 , fig. 13 (♂ terminalia). Type-locality: Brazil, São Paulo, Salesópolis, Estação Biológica de Boracéia. Holotype ♂, MZUSP.

Notes. The type series included the male holotype and seventeen paratypes without gender specification, of which four male paratypes, all from the type-locality and with the male terminalia on slide mountings, are at the NHM

(BMNH(E) # 257661-4).

M. cramptoni ( Coher) 1950b: 569 , fig. 5 (♂ terminalia). Type-locality: Brazil, São Paulo, Salesópolis, Estação Biológica de Boracéia. Holotype ♂, MZUSP.

Notes. The type series included the male holotype and fifteen paratypes without gender specification, of which two male paratypes, all from the type-locality and with the male terminalia in slide mountings, are at the NHM

(BMNH(E) # 257670-1).

M. ferrazi ( Coher) 1950b: 579 , fig. 15 (♂ terminalia). Type-locality: Brazil, São Paulo, Salesópolis, Estação Biológica de Boracéia. Holotype ♂, MZUSP.

Notes. The type series included the male holotype and fifteen paratypes without gender specification, of which three male paratypes from the type-locality (with male terminalia on slide mountings) are at the NHM ( BMNH (E)

# 257665-7).

M. hansoni ( Coher) 1950b: 566 , fig. 3 (♂ terminalia). Type-locality: Brazil, São Paulo, Salesópolis, Estação Biológica de Boracéia. Holotype ♂, MZUSP.

Notes. The type series included the male holotype and thirty-one paratypes without gender specification, of which four male paratypes from the type-locality (with male terminalia preparations) are at the NHM ( BMNH (E) #

257644-7).

M. lanei ( Coher) 1950b: 572 , fig. 9 (♂ terminalia). Type-locality: Brazil, São Paulo, Salesópolis, Estação Biológica de Boracéia. Holotype ♂, MZUSP.

Notes. The type series included the male holotype and eighteen paratypes without gender specification, of which three male paratypes from the type-locality (with male terminalia preparations) are at the NHM ( BMNH (E) #

257672-4).

M. manteri ( Coher) 1950b: 564 , fig. 1 (♂ terminalia). Type-locality: Brazil, São Paulo, Salesópolis, Estação Biológica de Boracéia. Holotype ♂, MZUSP.

Notes. The type series included the male holotype and thirty-two paratypes without gender specification, of which six male paratypes are present at the NHM, all coming from the type-locality (with male terminalia preparations)

(BMNH(E) # 257636-40/55).

M. mathesoni ( Coher) 1950b: 578 , fig. 14 (♂ terminalia). Type-locality: Brazil, São Paulo, Salesópolis, Estação Biológica de Boracéia. Holotype ♂, MZUSP.

Notes. The type series included the male holotype and fifteen paratypes without gender specification, of which six male paratypes from the type-locality (with male terminalia preparations) are at the NHM ( BMNH (E) # 257641-3) .

M. samesteri ( Coher) 1950b: 576 , fig. 12 (♂ terminalia). Type-locality: Brazil, São Paulo, Salesópolis, Estação Biológica de Boracéia. Holotype ♂, MZUSP.

Notes. The type series included the male holotype and twenty-five paratypes without gender specification, of which four male paratypes from the type-locality (with male terminalia preparations) are at the NHM ( BMNH (E) #

257657-60).

M. shannoni ( Coher) 1950b: 574 , fig. 10 (♂ terminalia). Type-locality: Brazil, São Paulo, Salesópolis, Estação Biológica de Boracéia. Holotype ♂, MZUSP.

Notes. The type series included the male holotype and twelve paratypes without gender specification, of which two male paratypes from the type-locality and with male terminalia preparations are at the NHM ( BMNH (E) # 257668-

9).

Unplaced Mycomya species

M. coxalis Freeman 1951: 43 , figs. 57–58 (♂ terminalia). Type-locality: Argentina, Río Negro, Bariloche. Holotype ♂, NHM.

( Figure 157 )

Notes. The type series included the male holotype, fourteen male and one female paratypes, of which twelve male and two female paratypes are at the NHM , so it is probable that Freeman misidentified the gender of one of the paratypes. The holotype labels are: “ Mycomya / bifida [hw] / Freeman. / HOLOTYPE ” [printed on red label]” // [male terminalia on a microslide attached to an acetate sheet] // “HOLO- / TYPE” [printed in black on a rounded label with red border]” // “Bariloche / 16 [hw]. xi.1926.” // “ Argentina: / Terr. Río Negro. / F. & M. Edwards / B.M. 1927–63.” // “BMNH(E) # / 257598”. The holotype is in excellent condition. No paratypes have the terminalia dissected ( BMNH (E) # 257599-612) .

M. cylindrica Freeman 1951: 41 , figs. 50–52 (♂ terminalia). Type-locality: Chile, Llanquihue , Casa Pangue. Holotype ♂, NHM.

( Figure 158 )

Notes. The type series included the male holotype, a male and a female paratypes, all of which are at the NHM . The holotype labels are: “ Mycomya / bifida [hw] / Freeman. / HOLOTYPE ” [printed on red label]” // [male terminalia on a microslide attached to an acetate sheet] // “HOLO- / TYPE” [printed in black on a rounded label with red border]” // “Peulla / 12–13.xii.1926.” // “ Argentina: / Terr. Río Negro. / F. & M. Edwards / B.M. 1927– 63.” // “BMNH(E) # / 257559”. The holotype is in good condition. None of the paratypes have the terminalia dissected ( BMNH (E) # 257560-1) .

M. divisa Freeman 1951: 45 , figs. 68–70 (♂ terminalia). Type-locality: Chile, Llanquihue , Puerto Varas. Holotype ♂, NHM.

( Figure 159 )

Notes. The type series included the male holotype and six male paratypes, of which the holotype and five paratypes are at the NHM . The labels of the holotype are: “HOLO- / TYPE” [printed in black on a rounded label with red border]” // “ Mycomya / divisa [hw] / Freeman. / HOLOTYPE ” [printed on red label]” // [male terminalia dissected on an acetate sheet] // “Puerto Varas / 16.xii.1926.” // “South Chile / Llanquihue prov. / F. & M. Edwards / B.M. 1927–63.” // “BMNH(E) # / 257625”. The holotype is in good condition, only one of the legs is missing. One of the paratypes has the male terminalia dissected ( BMNH (E) # 257626-30) .

M. falcifera Freeman 1951: 41 , figs. 47–49 (♂ terminalia). Type-locality: Argentina, Rio Negro, Lago Correntoso. Holotype ♂, NHM.

( Figure 160 )

Notes. The type series included the male holotype, sixteen male and two female paratypes, of which the holotype, fourteen male and both female paratypes at the NHM. The holotype labels are: “ Mycomya / falcifera [hw] / Freeman. / HOLOTYPE ” [printed on red label]” // [male terminalia on a microslide attached to an acetate sheet] // “HOLO- / TYPE” [printed in black on a rounded label with red border]” // “Lago Correntoso. / 18–25.xi.1926.” // “ Argentina: / Terr. Río Negro. / F. & M. Edwards / B.M. 1927–63.” // “BMNH(E) # / 257543). The holotype is in excellent condition. One paratype had the terminalia dissected ( BMNH (E) # 257544-58). One of the specimens labeled as a paratype does not have a NHM type number label .

M. flavescens Freeman 1951: 38 , figs. 40–41 (♂ terminalia), 75 (midcoxal process). Type-locality: Argentina, Rio Negro, Lago Correntoso. Holotype ♂, NHM.

( Figure 161 )

Notes. The type series included only the male holotype. Its labels are: [male terminalia on a microslide attached to an acetate sheet] // “ Mycomya / flavescens [hw] / Freeman. / HOLOTYPE ” [printed on red label]” // [preparation of mid leg on a microslide attached to an acetate sheet] // “HOLO- / TYPE” [printed in black on a rounded label with red border]” // “Lago Correntoso. / 18–25.xi.1926.” // “ Argentina: / Terr. Río Negro. / F. & M. Edwards / B.M. 1927–63.” // “BMNH(E) # / 257527”. Only one of the legs is incomplete, otherwise the holotype is in good condition.

M. fuscicornis Freeman 1951: 36 , figs. 24–25 (♂ terminalia), 81 (midcoxal process). Type-locality: Chile, Chiloé , Puntra. Holotype ♂, NHM.

( Figure 162 )

Notes. The type series included only the male holotype. The labels with the holotype are: “ Mycomya / fuscicornis [hw] / Freeman. / HOLOTYPE ” [printed on red label]” // [male terminalia on a microslide attached to an acetate sheet] // “HOLO- / TYPE” [printed in black on a rounded label with red border]” // “Puntra / 19.xii.1926.” // “S. Chile: / Chiloé I. / F. & M. Edwards / B.M. 1927–63.” // “BMNH(E) # / 257474”. The abdomen without the terminalia is glued to the micropin base, together with one of the wings.

M. inermis Freeman 1951: 40 , figs. 44–46 (♂ terminalia). Type-locality: Argentina, Rio Negro, Lago Correntoso. Holotype ♂, NHM.

( Figure 163 )

Notes. The type series included the male holotype, ten male and five female paratypes, of which nine male and four female paratypes are at the NHM. The labels of the holotype are: “ Mycomya / inermis [hw] / Freeman. / HOLOTYPE ” [printed on red label]” // [male terminalia on a microslide attached to an acetate sheet] // “HOLO- / TYPE” [printed in black on a rounded label with red border]” // “Lago Correntoso. / 18–25.xi.1926.” // “ Argentina: / Terr. Río Negro. / F. & M. Edwards / B.M. 1927–63.” // “BMNH(E) # / 257529”. The holotype is in excellent condition. The paratypes at the NHM are numbered BMNH(E) # 257530-42. One of the paratypes has a preparation of the male terminalia .

M. infuscata Freeman 1951: 39 , figs. 42–43 (♂ terminalia). Type-locality: Chile, Angol. Holotype ♂, NHM.

( Figure 164 )

Notes. The type series included only the male holotype. The labels of the holotype are: “ Mycomya / infuscata [hw] / Freeman. / HOLOTYPE ” [printed on red label]” // [male terminalia on a microslide attached to an acetate sheet] / / “HOLO- / TYPE” [printed in black on a rounded label with red border]” // “AngolChile / D.S. Bullock” // “Angol Chile [pr] / 3.august.1929 [hw] / B.M. 1927–63.” // “BMNH(E) # / 257528”. The holotype is in fair condition, one wing entirely lost, the other wing missing its distal three fourth.

M. jaffuelensis Freeman 1951: 43 , figs. 59–60 (♂ terminalia). Type-locality: Chile, Valparaiso, Marga Marga. Holotype ♂, NHM.

( Figure 165 )

Notes. The type series included the male holotype, two male and two female paratypes, of which the holotype, a male and a female paratypes, and a specimen with the abdomen missing are at the NHM. The holotype labels are: “ Mycomya / jaffuelensis [hw] / Freeman. / HOLOTYPE ” [printed on red label]” // [male terminalia on a microslide attached to an acetate sheet] // “HOLO- / TYPE” [printed in black on a rounded label with red border]” // “ Chile / Marga-Marga / ix.1929. / F. Jaffuel / B.M. 1929–480.” // “BMNH(E) # / 257613”. The holotype is in fair condition. The antennae are both missing, as well as one mid- and one hindlegs, and most of the tarsi. The numbers of the three paratypes are BMNH(E) # 257614-6 .

M. lutea Enderlein 1910a: 161 . Type-locality: Brazil, Santa Catarina. Holotype ♂, PAN.

Notes. There are some identified specimens from Rio Claro and Juquiá (São Paulo). The identity of these specimens with the type is questionable .

M. meridionalis (Johannsen) 1909: 180 (nom. nov. for Sciophila diluta Williston ). Type-locality: St. Vincent, W.I.

Lectotype ♂, NHM (here designated).

M. DILUTA (Williston) 1896: 263, Pl. VIII, fig. 17 (wing) ( Sciophila ) (preocc. Zetterstedt 1860). Type-locality:

St. Vincent, W.I. Lectotype ♂, NHM (here designated).

Notes. The type series included two male syntypes, both of which are at the NHM, but they are apparently one male and one female, both from the type-locality. The specimens are in fair condition, the specimen choosen as lectotype with both wings and three legs missing, and the paralectotype with one wing, one fore- and one midleg missing. The more damaged specimen was choosen as the lectotype because it is clearly a male and the terminalia is well preserved, even though not in a slide mounting. The labels of the lectotype are: “SYN- / TYPE” [printed in black on a rounded label with light blue border] // “Winward side / St. Vincent, W.I. / H.H. Smith.” // “ 1000 feet ” / / “W. Indies. / 1907-66.” // “BMNH(E) # / 257370”. “ SYNTYPE of / Sciophila diluta / Williston, 1896 / det. N.P. Wyatt 2002” / “ Sciophila / diluta [hw] / Williston. [printed in red on a folded paper with red borders. The paralectotype number is BMNH(E) # 257371.

M. pectinata Freeman 1951: 43 , figs. 61–63 (♂ terminalia). Type-locality: Argentina, Rio Negro, Lago Correntoso. Holotype ♂, NHM.

( Figure 166 )

Notes. The type series included the male holotype, a male and a female paratype, all of which are at the NHM. The labels of the holotype are as follows: “ Mycomya / pectinata [hw] / Freeman. / HOLOTYPE ” [printed on red label]” // [male terminalia on a microslide attached to an acetate sheet] // “HOLO- / TYPE” [printed in black on a rounded label with red border]” // “Lago Correntoso. / 18–25.xi.1926.” // “ Argentina: / Terr. Río Negro. / F. & M. Edwards / B.M. 1927–63.” // “BMNH(E) # / 257617”. The holotype is in good condition. The paratype numbers are BMNH(E) # 257618-9 .

M. setifera Freeman 1951: 45 , figs. 71–74 (♂ terminalia). Type-locality: Chile, Chiloé , Castro. Holotype ♂, NHM. ( Figure 167 )

Notes. The type series included the male holotype, two male and one female paratypes, all of which are at the NHM. The holotype labels are: “ Mycomya / setifera [hw] / Freeman. / HOLOTYPE ” [printed on red label]” // [male terminalia on a microslide attached to an acetate sheet] // “HOLO- / TYPE” [printed in black on a rounded label with red border]” // “Castro. / 20–22.xi.1926.” // “S. Chile: / Chiloé I. / F. & M. Edwards / B.M. 1927–63.” // “BMNH(E) # / 257631”. The holotype is in fairly good condition. The paratypes are numbered BMNH(E) # 257632-4, none of which have the male terminalia dissected; the abdomen of probably a male paratype is entirely missing .

M. spinifera Freeman 1951: 36 , figs. 22–23 (♂ terminalia), 79 (midcoxal process). Type-locality: Chile, Chiloé , Ancud. Holotype ♂, NHM.

( Figure 168 )

Notes. The type series included the male holotype and two paratypes, all of which are at the NHM. The holotype labels are: “ Mycomya / spinifera [hw] / Freeman. / HOLOTYPE ” [printed on red label]” // [male terminalia on a microslide attached to an acetate sheet] // “HOLO- / TYPE” [printed in black on a rounded label with red border]” // “Ancud / 17–19.xii.1926.” // “S. Chile: / Chiloé I. / F. & M. Edwards / B.M. 1927–63.” // “BMNH(E) # / 257471”. The holotype is in fairly good condition, only one of the antennae broken. One of the paratypes ( BMNH (E) # 257472-3) has the terminalia dissected and mounted .

M. spinosa Freeman 1951: 36 , figs. 22–23 (♂ terminalia), 79 (midcoxal process). Type-locality: Chile, Chiloé , Puntra. Holotype ♂, NHM.

( Figure 169 )

Notes. The type series included the male holotype and a male paratype. The labels with the paratype are as follows: “ Mycomya / spinosa [hw] / Freeman. / HOLOTYPE ” [printed on red label]” // [male terminalia on a microslide attached to an acetate sheet] // “HOLO- / TYPE” [printed in black on a rounded label with red border]” // “Puntra / 19.xii.1926.” // “S. Chile: / Chiloé I. / F. & M. Edwards / B.M. 1927–63.” // [a small piece of paper with some handwritten notes (“abdomen blackish”) ( BMNH (E) # 257475). The holotype is in excellent condition. The paratype number is BMNH(E) # 257476.

M. taurus Freeman 1951: 44 , figs. 64–67. Type-locality: Argentina, Rio Negro, Lago Correntoso. Holotype ♂, NHM.

( Figure 170 )

Notes. The type series included the male holotype and six male paratypes, of which the holotype and four paratypes are at the NHM. The labels of the holotype are: “ Mycomya / taurus [hw] / Freeman. / HOLOTYPE ” [printed on red label]” // [male terminalia on a microslide attached to an acetate sheet] // “HOLO- / TYPE” [printed in black on a rounded label with red border]” // “Lago Correntoso. / 18–25.xi.1926.” // “ Argentina: / Terr. Río Negro / F. & M. Edwards / B.M. 1927–63.” // “BMNH(E) # / 257620”. The holotype is in fair condition. The part of the abdomen of the holotype was glued to the base of micropin and some of the legs are broken. The paratypes are numbered BMNH(E) # 257621-4. One of them is particularly damaged, with only a wing and some left legs, but there is a male paratype in excellent condition. No paratypes had their terminalia dissected .

M. trichops Freeman 1951: 35 , figs. 20–21 (♂ terminalia), 77 (midcoxal process). Type-locality: Chile, Chiloé , Ancud. Holotype ♂, NHM.

( Figure 171 )

Notes. The type series included the male holotype and a single male paratype, all of which are at the NHM. The labels of the holotype are: “ Mycomya / trichops [hw] / Freeman. / HOLOTYPE ” [printed on red label]” // [male terminalia on a microslide attached to an acetate sheet] // “HOLO- / TYPE” [printed in black on a rounded label with red border]” // “Ancud / 17–19.xii.1926.” // “S. Chile: / Chiloé I. / F. & M. Edwards / B.M. 1927–63.” // “BMNH(E) # / 257469”. The holotype is in fair condition, with both antennae distally broken and one of the wings partially folded. The paratype is numbered BMNH(E) # 257470 .

Figures

FIGURES 152–161. Mycetophilidae habitus. 152. Mycomya (Mycomya) ochracea Freeman, holotype ♂. 153. Mycomya (Mycomya) simillima Freeman, holotype ♂. 154. Mycomya (Mycomya) subfusca Freeman, holotype ♂. 155. Mycomya (Mycomyopsis) forcipata Freeman, holotype ♂. 156. Mycomya (Mycomyopsis) longistila Freeman, holotype ♂. 157. Mycomya coxalis Freeman, holotype ♂. 158. Mycomya cylindrica Freeman, holotype ♂. 159. Mycomya divisa Freeman, holotype ♂. 160. Mycomya falcigera Freeman, holotype ♂. 161. Mycomya flavescens Freeman, holotype ♂.

FIGURES 162–174. Mycetophilidae habitus. 162. Mycomya fuscicornis Freeman, holotype ♂. 163. Mycomya inermis Freeman, holotype ♂. 164. Mycomya infuscata Freeman, holotype ♂. 165. Mycomya jaffuelensis Freeman, holotype ♂. 166. Mycomya pectinata Freeman, holotype ♂. 167. Mycomya setifera Freeman, holotype ♂. 168. Mycomya spinifera Freeman, holotype ♂. 169. Mycomya spinosa Freeman, holotype ♂. 170. Mycomya taurus Freeman, paratype ♂. 171. Mycomya trichops Freeman, holotype ♂. 172. Neoempheria biflagellata Edwards, holotype ♂. 173. Neoempheria bilobata Edwards, holotype ♂. 174. Neoempheria bipectinata Edwards, paratype ♀.