Monoctenus fujisanus Togashi, 2001

Hara, Hideho & Nagase, Hirohiko, 2023, The Japanese species of Monoctenus (Hymenoptera: Diprionidae), Zootaxa 5380 (6), pp. 541-561 : 548-550

publication ID	https://doi.org/ 10.11646/zootaxa.5380.6.3
publication LSID	lsid:zoobank.org:pub:AA915DFA-58B9-428D-AF5D-193E3295AB04
DOI	https://doi.org/10.5281/zenodo.10259700
persistent identifier	https://treatment.plazi.org/id/03B38786-FF93-CD57-FF21-F8C1FA7A52E2
treatment provided by	Plazi
scientific name	Monoctenus fujisanus Togashi, 2001
status

Treatment

( Figs 5A View FIGURE 5 2b View FIGURE 2 , 6A 2b, B2b View FIGURE 2 , 7A 2, B2 View FIGURE 2 , 14)

Monoctenus fujisanus Togashi, 2001: 44 ; Taeger et al. 2010: 211; Yoshida 2010: 28; Hara 2019: 46; Hara 2020: 287. Monoctenus itoi : Togashi 2001: 41 (part, female); Hara 2020: 287 (part, female). Not Okutani, 1958.

Additional description. Female. Length 7.5–8.7 mm. Figs 5A View FIGURE 5 2b View FIGURE 2 , 14A, B. Head capsule black, with supraclypeal area yellow to red brown, anterior tentorial pit and its surroundings usually red yellow to red brown; clypeus dorsomedially dark brown, dorsally widely red yellow or almost entirely red yellow; vertex posteromedially dark brown or posteriorly widely red yellow; frons sometimes anterolaterally red yellow ( Fig. 14C–J View FIGURE 14 ). Labrum brown to black. Prepectus red yellow or black. Postspiracular sclerite black. Mesoscutum black on posterolateral sunken part and often on lateral part of lateral lobe ( Fig. 14L–N View FIGURE 14 ). Mesoscutellum sometimes narrowly red yellow anteriorly. Mesepisternum black ( Fig. 14B View FIGURE 14 ), sometimes with pectus red yellow ( Fig. 5A View FIGURE 5 2b View FIGURE 2 ). Abdomen without pale parts ( Fig. 14A, B View FIGURE 14 ).

Clypeus without median furrow ( Fig. 14G–J View FIGURE 14 ); ventral edge widely concave. Antenna with 15–17 antennomeres ( Figs 7A View FIGURE 7 2 View FIGURE 2 , 14K); flagellomere 6 in lateral view with breadth including serration 1.1–1.5 × dorsal length. Valvula 3 ( Fig. 14O–Q View FIGURE 14 ) in dorsal view tapering and with apex narrowly rounded or pointed, in lateral view with apex narrowly rounded and dorsal edge nearly straight. Lance in lateral view with dorsal edge gently rounded near middle and almost straight from middle to near apex ( Fig. 14R View FIGURE 14 ); radix long. Lancet with 9–10 annuli ( Figs 7B View FIGURE 7 2 View FIGURE 2 , 14S, T); length from ventral end of ctenidium 1 to apex 4.4–5.3 × breadth; ctenidia 1 and 2 parallel, inclined posteriorly ( Fig. 7B View FIGURE 7 2 View FIGURE 2 ); ctenidia 3 and 4 converging dorsally; ctenidia 4–7 parallel, inclined anteriorly.

Male. Unknown.

Material examined. Holotype ( Figs 14A–C, G, K, M, O–Q, S View FIGURE 14 ): ♀, labelled “ Tenshokyo, 1000m, Mt. Fuji , Shizuoka Pref., 20. VI. 1989, H. Ishikawa”, “DIP-01” and “Holotype Monoctenus fujisanus n. sp. ” . Other material examined: 1♀, Tochigi Pref., Otawara, Shimoishigami, 18. V. 2010, E. Katayama ( Figs 5A View FIGURE 5 2b View FIGURE 2 , 6A 2b View FIGURE 2 ) (deposited in the collector’s collection) ; 1♀, Tochigi Pref., Yaita, Happogahara , 21. VI. 1987, T. Saito (cited as M. itoi by Hara 2020) ; 1♀, Kanagawa Pref., Hakone, Mt. Komagatake , 5. VI. 2004, H. Nagase (deposited in Kanagawa Prefectural Museum of Natural History, Odawara) ; 1♀, Yamanashi Pref., Nambu, Toshima , 5. V. 1977, A. Shinohara ( Figs 7A View FIGURE 7 2 View FIGURE 2 , 14F, J, N, T) (treated as M. itoi by Togashi 2001) ; 1♀, Nara Pref., Yoshino , 2. VI. 1916, Takeuchi ( Fig. 14E, I View FIGURE 14 ) ; 1♀, Hyogo Pref., Sasayama , 200m, 15. V. 1983, T. Naito ( Figs 6B View FIGURE 6 2b View FIGURE 2 , 7B 2 View FIGURE 2 , 14D, H, L, R) .

Distribution. Japan (Honshu) ( Togashi 2001).

Host plant. Unknown.

Remarks. This species is separated from other Monoctenus species as stated in the key and the remarks of M. cryptomeriae .

Togashi (2001) described M. fujisanus as a closely allied species to M. decoratus and separated their females by the differences in the angle of the front edge of a scutellum, the length ratio of an antennomere 3 to an antennomere 2, the number and shape of the annuli of a lancet, the length of a female cercus and the shape of a clypeus. However, these character states except for the shape of a lancet are variable in our material and not useful to distinguish them.

This species has considerable variation in the coloration of the female head and mesoscutum ( Fig. 14C–J, L–N View FIGURE 14 ). Togashi (2001) and Hara (2020) regarded the richly light-marked females as M. itoi . However, the females with different colorations do not have significant morphological differences, and their color differences are continuous.