Tasata chiloensis, RAMÍREZ, 2003

RAMÍREZ, MARTÍN J., 2003, The Spider Subfamily Amaurobioidinae (Araneae, Anyphaenidae): A Phylogenetic Revision At The Generic Level, Bulletin of the American Museum of Natural History 2003 (277), pp. 1-262 : 237-239

Tasata chiloensis , new species Figures 131 , 132

TYPES: Female holotype and male paratype from Chile, Región X, Chiloé province, Cordillera de San Pedro, Piroquina, 500 m, 10– 11.III.1987, L. Peña, deposited in AMNH.

ETYMOLOGY: The specific name refers to the type locality.

DIAGNOSIS: Distinguished by having semicircular anterior epigynal ridges, combined with a wide, posteriorly displaced anterior pouch (fig. 132F) and a transversely striated embolar base (fig. 132E).

FEMALE (holotype, fig. 132A): Total length 7.71. Carapace length 3.13, width 2.43, wid­ er on legs II–III. Length of tibia/metatarsus: I, 3.86/2.93; II, 3.59/2.67; III, 2.17/1.83; IV, 2.70/2.50. Palpal tarsus length 1.20. Chelicerae with two teeth on retromargin. Sternum length 1.57, width 1.20. Spines: leg I, femur d 1–1–1, p 0­ 0­ d1­ 1­ d1, r 0­ d1­ d1; tibia v 2– 2–2, p and r 1–1; metatarsus v 2bas, p and r 1–0, d 2ap. II = I. III, femur d 1–1–1, p and r 0­ d1­ d1; tibia v p1–2–2, p and r d1–1, d r1–1; metatarsus v 2–0­ p1 and group of apical hairs, p and r 0–1–1, d 0­ p1­ r1­ 2. IV, femur d 1–1–1, p and r d1ap; patella r 1; tibia = III; metatarsus v 2­ p1­ p1 and group of apical hairs, p and r d1–1–1, d 0­ p1–2. Abdomen length 4.52, width 2.00, spiracle–epigastrium 2.07, spiracle–spinnerets 0.80. Col­ or: yellow with small brown dots on legs I and II, patellae III and IV with dorsal apical spot. Dorsum with pattern of darker dots, abdomen with white guanine reticulum on sides, and ventral median dark band from epigastrium to tracheal spiracle. Epigyne (fig. 132F, G): lateral lobes separate, median field rugose, slightly elevated at sides of anterior pouch. Anterior pouch wide, surrounded by U­shaped suture, posterior border reaching epigastric furrow. Margins of lateral lobes continued into semicircular anterior carinae. Copulatory ducts surrounding spermathecae.

MALE (paratype): Total length 7.71. Carapace length 3.33, width 2.60. Length of tibia/ metatarsus: I, 6.12/4.92; II, 5.65/4.26; III, 3.46/2.60; IV, 4.12/3.46. Chelicerae as those of the female, right retromargin with three teeth (apical one smaller). Sternum length 1.63, width 1.27. Spines as in female, except: leg I, tibia d r1­ 0­ 1­ 0. II, tibia = I. III, femur p 0­ d1­ 1­ d1; patella r d1; metatarsus d 0­ p1– 2. IV, metatarsus p 0–1–1. Abdomen length 4.52, width 1.85, spiracle–epigastrium 2.03, spiracle–spinnerets 0.67. Color as in female. Palp (fig. 132B–E): tibia width/length 0.75, cymbium relatively large. Sperm duct with loop approximating anterior ventral margin of tegulum, which projects over paramedian apophysis. Embolus very thin, base with transverse anterior striae (fig. 132E), basal process absent, membranous area prolonged in lobe. Median apophysis long. Paramedian apophysis with retrolateral curved tip, one ventral/prolateral shorter projection. Primary conductor absent, only sclerotized stripe remains. Secondary conductor with two lateral prongs, one at each side of end of canal; prolateral portion with rounded lobe fitting un­ der embolus; canal ending in pointed projection, with prolateral, rugose prong; retrolateral portion large, weakly sclerotized, membranous at apical/retrolateral side, with basal elevated prong.

VARIABILITY: The three cheliceral teeth on right retromargin of paratype are abnormal. Spines: metatarsus IV v 2–2­ p1, p 0–1–1.

NATURAL HISTORY: This species builds retreats on foliage, most commonly on ‘‘colihue’’ bamboos ( Chusquea spp. , fig. 131A, B).

DISTRIBUTION: Humid forests in Chile, in Osorno, Llanquihue and Chiloé provinces.

OTHER MATERIAL EXAMINED: ARGENTINA: Neuquén: Puerto Blest, 7–20.I.2000, L. Lopardo and A. Quaglino, 2 immatures ( MACN­Ar). CHILE: Región X (Los Lagos): Osorno: El Mirador , 45 km W La Unión, 900 m, 1–2.III.1987, L. Peña, 43 3♀ ( AMNH) ; Puyehue Natl. Park : 700 m, 9.XII.1994, L. Peña, 1♀ ( AMNH) ; 480 m, 40°44̍S, 72°18̍W, 21.XI.1993, N. Platnick, K. Catley, M. Ramírez, T. Allen , 2♀ ( AMNH, eggsac in MACN­Ar, photo MJR 1411–1413), 1♀, 1 eggsac, 1 immature ( MACN­Ar) ; 12 km SE Aguas Calientes , 700 m, 21.XI.1993, N. Platnick, K. Catley, M. Ramírez, T. Allen , 1♀ ( AMNH) ; Aguas Calientes, 13–17.XII.1998, M. Ramírez, L. Compagnucci, C. Grismado, L. Lopardo, 4 immatures ( MACN­Ar). Llanquihue: Alerce Andino Natl. Park , elev. 100 m, 41°35̍S, 72°41̍S, 23.XI.1993, N. Platnick, K. Catley, M. Ramírez, T. Allen , 1♀ ( AMNH) .

Figures

Fig. 131. Tasata chiloensis, n. sp. (Osorno, Puyehue, photos MJR 1411, 1413). A. Retreat on bamboo. B. Same, female guarding eggsac.

Fig. 132. Tasata chiloensis, n. sp., female holotype, male from Osorno, El Mirador. A. Female. B. Male palp, ventral view. C. Same, retrolateral view. D. Male copulatory bulb, retrolateral view. E. Same, apical view. F. Epigyne, ventral view. G. Same, cleared. Scale bars = A, 1 mm; B, C, 0.5 mm; D–G, 0.2 mm.