Corasoides angusi, Humphrey, 2017

Humphrey, Margaret, 2017, A Revision and Cladistic Analysis of the Genus Corasoides Butler (Araneae: Desidae) with Descriptions of Nine New Species, Records of the Australian Museum 69 (1), pp. 15-64 : 30-33

Corasoides angusi sp. nov.

Figs 13 , 14

Holotype ♂, Kuper Ranges , 30 km NNE of Wau, PNG [ Papua New Guinea], 7°05'S 146°45'E, 31 Oct 1996, M. Humphrey, M. Moulds, W. Angus, KS.71659 ( AM) . Paratypes as follows: 1♀, same data as holotype, KS.71660 ; 10♀♀, same data, KS.71662 ; 1♂, same data except collected Nov 1996, KS.71661 ( AM) .

Other material examined. Body parts remaining from dissections and electrophoretic work from identified specimens, namely 1 pair male palps, 1 epigyne and pair chelicerae from female, data as for holotype ( AM).

Diagnosis. This species differs in the male from the closely related C. nimbus sp. nov. by having a strongly hooked retrolateral apophysis tapering to a fine point rather than a blunt, truncated hook and by the base of the embolus originating prolaterally rather than distally or retrolaterally. The female is distinguished by having only three bends (weakly convoluted) in the insemination ducts rather than at least five (strongly convoluted) in C. nimbus sp. nov. and C. stellaris sp. nov. Females of C. angusi sp. nov. cannot be readily distinguished externally from those of C. nimbus sp. nov.. However, the epigynes of both C. angusi sp. nov. and C. nimbus sp. nov. differ from C. stellaris sp. nov. in having the lateral extension of the scape no more than half the width of the epigyne.

Description. Medium to large spiders. Carapace. Full pattern for genus but indistinct in some specimens. Abdomen. Distinct basic abdominal pattern.

Male ( Figs 13a–f , 14b,d,f ). CL 6.3 (6.5), CW 4.3 (4.5), AL 6.6, AW 3.2, HW 3.1, EpGW 1.9, MOQL 0.85, MOQAW 0.65, MOQPW 0.85, SL 2.9, SW 2.7, ML 2.4, MW 1.3, LL 1.3, LW 0.9, ChelL 4.4 (4.5), ChelW 1.6 (1.6), clypeus height 0.5. Cephalothorax. Sternum roughly equal in length and width, distinct posterior point, long ventrally projecting hairs. Labium longer than wide, notched basally. Cheliceral teeth. retromarginal 6(7), promarginal 4, subequally spaced, no fusions. No transverse ridges between teeth margins. Metatarsus IV particularly long, other leg lengths:

I II III IV Palp femur 11.9 9.3 9.4 12.1 4.5

patella 2.8 2.5 1.9 2.3 1.3

tibia 12.6 8.4 6.8 10.0 2.5 metatarsus 14.9 12.0 11.5 16.3 —

tarsus 6.1 4.1 3.9 5.1 4.0

total 48.1 36.3 33.4 45.8 12.3

Spination. Leg I: femur d3,3,1,1,1,1,1,1,3,1,3; tibia d3,3,3,3,3,1 v1,1,1,1,1,1,1,2; metatarsus d2,1,1,1,1,2 v2,2,2,3. Leg II: femur d3,3,2,3,1,3,3; tibia 1,1,1,1,1,2 v1,1,1,1,1,1,1,1,2; metatarsus d1,1,1,1,1,1,2v2,2,2,2,3. Leg III: femur d3,2,1,3,3,2,2; tibia d1,1 v1,1,1,1,1,1,1,1,2; metatarsus d2,2,2,2,2,2, v2,2,2,2. Leg IV: femur d2,2,3,3,1; tibia d1,1,1, v1,1,2,1,1,1; metatarsus d2,2,2,2,2,2,2,2, v1,2,2,1,1,1,1,2,2. Palp: femur d1,3,v1; tibia d1; tarsus several. Male palp. Digitiform portion of the cymbium very long- cymbium length about six times diameter of bulb. Retroventral apophysis absent. Ventral apophysis lobe-like, not enclosing any extension of the intersegmental membrane. Retrolateral apophysis long, hook-like, tapering and curving initially towards bulb and then pointing towards its base. Embolus arises prolaterally, enters conductor prolaterally. Conductor stalked, wide basally. Conductor tip pointed, twisted but with less than one rotation. Conductor tip without ridges and pointing towards cymbium, i.e. dorsally ( Fig. 13a–c ). Trichobothria on cymbium: single row of at least six evenly spaced on prolateral to dorsal line, decreasing in length proximally. Abdomen. Anterior lateral spinnerets with tail of small spigots.

Female ( Fig. 14a,c,e,f ), similar to male. CL 6.3 (5.6–7.0), CW 4.8 (4.6–4.9), AL 7.6, AW 5.1, HW 3.3, EpGW 1.8, MOQL 0.91, MOQAW 0.70, MOQPW 0.85, SL 3.3, SW 2.7, ML 2.5, MW 1.3, LL 1.4, LW 1.0, ChelL 3.6 (3.2–3.7), ChelW 1.6, clypeus height 0.4. Cheliceral teeth as for male. Leg lengths:

I II III IV Palp femur 8.8 7.3 6.0 7.5 4.0

patella 2.5 2.4 2.0 2.3 1.3

tibia 9.0 6.4 4.3 6.9 3.5 metatarsus 9.0 7.1 7.0 9.0 —

tibia 3.8 2.9 2.4 3.0 3.3

total 29.3 26.0 21.6 28.6 12.0

Spination. Leg I: femur d2,1,2,1,1,2,4; tibia; metatarsus d2 v2,1,2,2,1,1,2. Leg II femur d3,1,3,2,1,1,2,3; tibia d1,1,2 v1,1,2,2. Leg III: femur d3,3,1,2,1,2,4; tibia d1,1,1,3 v2,1,1,1; metatarsus; d2,1,1,1,1,2 v1,2,2,2. Leg IV: femur d1,2,1,1,2,1,2,3; tibia d2,2,2,2 v2,2,1,1,1; metatarsus; d1,1,1,1,1,2,4 v1,1,1,1; Palp; femur; patella d1,1, tibia d1,1, p1; tarsus several. Trichobothria on first tarsus: at least eight, evenly spaced, decreasing in length proximally. Epigyne ( Fig. 14a ). Height 0.8, width 1.3. Smooth, no ridges or beaks. Genital openings on transverse midline. Scape stalk short, almost equal in width to diameter of a genital opening. Lateral extensions of scape wide and extending almost to lateral margins of genital openings. Anterior and lateral surfaces of epigyne covered in long, posteriorly directed hairs. Insemination ducts, weakly convoluted, proceed anteriorly from genital openings with three bends, small diverticula at site of entry into spermathecae ( Fig. 14c,e ).

Habitat. High altitude rainforest, in particular, mist forest. Webs with retreat into crevices in tree trunks, earthen embankments and fallen logs, up to 1.5 m from ground level.

Distribution. Common in Kuper Ranges, 30 km NNE of Wau, PNG, 7°05'S 146°45'E, Papua New Guinea ( Fig. 14f ).

Etymology. Named after Warrick Angus who enthusiastically helped to collect the type series.

Figures

Figure 13. Corasoides angusi sp. nov., male: (a) palpal organs, prolateral; (b) palpal organs, ventral; (c) palpal organs, retrolateral; (d) face; (e) cephalothorax, lateral; (f) sternum, labium and maxillae, dorsal.

Figure 14. Corasoides angusi sp. nov.: (a) female epigyne; (b) male