Tityus (Archaeotityus), 2006

Moreno-González, Jairo A., O, Ranulfo González & D, Eduardo Flórez, 2019, Taxonomic revision of the Colombian Tityus (Archaeotityus) (Scorpiones, Buthidae) species: a morphological and morphometric approach, with a description of a new species, Zootaxa 4660 (1), pp. 1-94 : 82-83

publication ID	https://doi.org/ 10.11646/zootaxa.4660.1.1
publication LSID	lsid:zoobank.org:pub:DA70C158-7FB0-4B46-9A1D-EA32D24B19B6
DOI	https://doi.org/10.5281/zenodo.4648606
persistent identifier	https://treatment.plazi.org/id/03B387E6-320D-1066-FF3E-F8D4DDD53F89
treatment provided by	Plazi
scientific name	Tityus (Archaeotityus)
status

Treatment

Identification key to the Colombian species of Tityus (Archaeotityus)

1. Subaculear tubercle large-sized pyramidal, and crest-like, with a blunt apex in lateral view ( Figures 19A, C, E View FIGURE 19 , 20A, C, E View FIGURE 20 ); metasomal segment V with three pairs of VSM macrosetae (VSM1, VSM2 and VSM3) ( Figures 17A, C, E View FIGURE 17 , 18A, C, E View FIGURE 18 ).... 2

1’. Subaculear tubercle medium-sized pyramidal, and triangle-like, with an acute and coarse apex in lateral view ( Figures 19B, D View FIGURE 19 , 20B, D View FIGURE 20 ); metasomal segment V with two pairs (VSM1 and VSM3) ( Figures 17D View FIGURE 17 , 18D View FIGURE 18 ) or three pairs (VSM1, VSM2 and VSM3) of VSM macrosetae ( Figures 17B View FIGURE 17 , 18B View FIGURE 18 )............................................................. 4

2. Metasomal segments II–IV without conspicuous distoterminal granules on the DL carinae ( Figure 40C, D View FIGURE 40 ); metasomal segment V with the VSM2 macrosetae proximal to the level of the VL2 macrosetae ( Figures 17E View FIGURE 17 , 18E View FIGURE 18 ).............. Tityus tayrona View in CoL

2’. Metasomal segments II–IV each has a conspicous distoterminal granule on the DL carinae ( Figures 15A, C View FIGURE 15 , 28A, B View FIGURE 28 ); metasomal segment V with the VSM2 macrosetae slightly distal or at the same level of the VL2 macrosetae ( Figures 17A, C View FIGURE 17 , 18A, C View FIGURE 18 )................................................................................................. 3

3. General coloration pattern with a dark yellow background moderately covered with dark brown spots ( Figure 24 View FIGURE 24 ); metasomal segment V completely dark reddish-brown with a few ventral yellow spots and a dorsomedian area without spots ( Figure 6 View FIGURE 6 C–F); metasomal segments with an acute and small (II–III) or a coarse and blunt (IV) distoterminal granule on the DL carinae ( Figures 15C View FIGURE 15 , 28A, B View FIGURE 28 )................................................................. Tityus guane View in CoL sp. nov.

3’. General coloration pattern with a dark yellow background densely covered with dark reddish-brown spots ( Figure 4 View FIGURE 4 ); metasomal segment V completely black ( Figure 6A, D View FIGURE 6 ); metasomal segments II–IV each has a medium-sized acute distoterminal granule on the DL carinae ( Figures 15A View FIGURE 15 , 16A, B View FIGURE 16 ).................................................. Tityus bastosi View in CoL

4. Metasomal segments with an acute and small (II–III) or a blunt and coarse (IV) distoterminal granule on the DL carinae ( Figures 15B View FIGURE 15 , 16C, D View FIGURE 16 ); metasomal segment V with the VSM2 macrosetae slightly distal or at the same level of VL2 ( Figures 17B View FIGURE 17 , 18B View FIGURE 18 ); telson VM carina composed of coarse granules and strongly crenulate in lateral view ( Figures 19B View FIGURE 19 , 20B View FIGURE 20 )................................................................................................. Tityus columbianus View in CoL

4’. Metasomal segments II–IV without conspicuous distoterminal granules on the DL carinae ( Figures 39A View FIGURE 39 . 40A, B View FIGURE 40 ); metasomal segment V without the VSM2 macrosetae ( Figures 17D View FIGURE 17 , 18D View FIGURE 18 ); telson VM carina composed of fine granules and feebly crenulate in lateral view ( Figures 19D View FIGURE 19 , 20D View FIGURE 20 )......................................................... Tityus parvulus View in CoL