Trimeresurus (Parias) schultzei (Griffin, 1909)

Leviton, Alan E., Siler, Cameron D., Weinell, Jeffrey L. & Brown, Rafe M., 2018, Synopsis of the Snakes of the Philippines, Proceedings of the California Academy of Sciences 64 (14), pp. 399-568 : 471-507

publication ID

https://doi.org/ 10.5281/zenodo.11512589

persistent identifier

https://treatment.plazi.org/id/03B38B5C-843D-FFAA-FFDC-FD9CFD75FA3C

treatment provided by

Felipe

scientific name

Trimeresurus (Parias) schultzei (Griffin, 1909)
status

 

Trimeresurus (Parias) schultzei (Griffin, 1909) View in CoL

Schultz’s Philippine Pitviper

Trimeresurus schultzei Griffin, 1909:601 View in CoL ; 1911:267.— Taylor, 1922a:292, pl. 36.— Leviton, 1964c:263.— Toriba, 1993a:105.— David & Ineich, 1999:290.— Gumprecht, Tillack, Orlov, Captain, and Ryabov, 2004:37, 9 col. photos (pp. 288–289). Trimeresurus (Parias) schultzei, Leviton, Brown, and Siler, 2014:518 View in CoL , fig. 13.

TYPE LOCALITY AND TYPE SPECIMEN (S).— Palawan [Iwahig]. Holotype: PNM 315 View Materials (destroyed during WW II).

PHILIPPINE DISTRIBUTION (ENDEMIC) ( Map 36B [p. 147]).— Balabac, Palawan.

CONSERVATION STATUS [IUCN].— Least Concern [2016] ver. 3.1.

Tropidolaemus philippensis ( Gray, 1842) Photo figure 114 Philippine Temple Pitviper

Trimeresurus philippensis Gray, 1842:48 View in CoL .— Taylor, 1922a:295, pl. 37, fig.1.

Tropidolaemus hombronii Guichenot in Jacquinot and Guichenot [q.v.], 1853:23, pl. 2, fig. 3.

Trimeresurus wagleri, Leviton, 1964c:265 View in CoL .

Tropidolaemus wagleri, Toriba, 1993b View in CoL , 108.— David & Ineich, 1999:295 (doubtfully included in T. wagleri View in CoL ).

Tropidolaemus philippensis, Vogel, David, Lutz, van Rooijen, and Vidal, 2007:31 View in CoL View Cited Treatment , figs. 25–26.— Leviton, Brown, and Siler, 2014:518, figs. 52B, E–F.— Wallach, Williams, and Boundy, 2014:734.— Sanguila, Cobb, Siler, Diesmos Alcala, and Brown, 2016:119, fig. 80.

TYPE LOCALITY AND TYPE SPECIMEN (S).— Philippines. Holotype: BMNH 1946.1 .17.67 (fide Vogel et al. [2007:31]).

PHILIPPINE DISTRIBUTION (ENDEMIC) ( Map 36C [p. 147]).— Dinagat, Leyte, Mindanao (Prov.: Agusan del Norte, Bukidnon, Cotabato, Davao City, Davao del Norte, Misamis Oriental), Samar.

REMARKS.— Tropidolaemus philippensis and T. subannulatus were formerly included in the ubiquitous species T. wagleri , but the assignment was seriously questioned by David and Ineich (1999:295–296). Indeed, recent studies have demonstrated that several recognizeable species have been inappropriately parading under the nominal taxon T wagleri , T. philippensis among them, as well as populations known from southern Mindanao and the Zamboanga Peninsula. On the large and topographically diverse island of Mindanao, and on careful examination, the Zamboanga population appears to be quite distinct from populations inhabiting other parts of the island, although Vogel et al. (2007) referred it to T. philippensis . Recent unpublished studies by Brown et al. suggest otherwise, and it is likely that the taxon T. hombronii (type locality, Zamboanga) will have to be resurrected from the synonymy of T. philippensis to accommodate the Zamboanga and, possibly, the Basilan populations. Furthermore, recently documented populations of tropidolaemids found on Leyte, Dinagat, Samar, and northeast Mindanao bear strong resemblances to T. philippensis but also to T. subannulatus (see Vogel et al. [2007:30, fig. 24] from Negros; also figs. 79–80 in Sanguila et al. [2016]). We have refrained from suggesting any changes at this time because this too is said to be under study by Vogel and David (see also remarks under Trimeresurus [ Parias ] flavomaculatus and Tropidolaemus subannulatus ). For additional details, see Leviton, Brown, and Siler (2014:518 and figs. 52B, E–F).

CONSERVATION STATUS [IUCN].— The conservation status of Tropidolaemus philippensis has not been assessed for the IUCN Red List [2016] ver. 3.1, but IUCN notes that it is listed in the Catalogue of Life.

Tropidolaemus subannulatus ( Gray, 1842) Photo figures 115–119 Philippine Temple Pitviper; Northern Philippine Temple Pit Viper; Bornean Keeled Green Pit Viper

Trimesurus subannualtus Gray, 1842:48 .

Trimeresurus wagleri View in CoL (part), Taylor, 1922a:296.— Leviton, 1964c:265.— Toriba, 1993b:108.— David & Ineich, 1999:295.

Trimeresurus wagleri alboviridis Taylor, 1917:366 (type locality Isabela, Occidental Negros, Negros Id., Philippines; holotype: CM R2433) View in CoL .

Trimeresurus wagleri wagleri, Taylor, 1922c:302 View in CoL .

Tropidolaemus wagleri, Burger, 1971:2 View in CoL , 20, 41, 86 (part).— Gaulke, 1994b:141 (part); 1996:51, figs. 5–6.— McDiarmid, Campbell, and Touré, 1999:349 (part).— Gumprecht, Tillack, Orlov, Captain, and Ryabov, 2004:41–42 (part), 8 col. photos (pp. 339–340).

Tropidolaemus cf. wagleri, Ferner, Brown, Sison, and Kennedy, 2001:55 View in CoL [22], figs. 51–52.

Tropidolaemus subannulatus, Vogel, David, Lutz, van Rooijen, and Vidal, 2007:23 View in CoL View Cited Treatment , figs. 12–16.— Gaulke, 2011:340–341, figs. 234–236.— McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra, and Brown, 2011:190.— Brown, Siler, Oliveros, Welton, Rock, Swab, Van Weerd, van Beijnen, Jose, Rodriguez, Jose, and Diesmos, 2013:93.— Leviton, Brown, and Siler, 2014:519, figs. 5A–B, 11, 12A–C, 52A, C–D.— Sanguila, Cobb, Siler, Diesmos Alcala, and Brown, 2016:109, fig. 79.

TYPE LOCALITY AND TYPE SPECIMEN (S).— Philippines. Syntypes (2): BMNH 1946.1.19.32–33 (formerly BMNH i.2.7a).

PHILIPPINE DISTRIBUTION ( Map 36D [p. 147]).— Balabac; Basilan, Bohol, Dinagat, Leyte, Luzon (Prov.: Albay, Bulacan, Cavite, Cagayan, Camarines Norte, Isabela, Laguna, Quezon), Mindanao (Prov.: Agusan del Norte, Agusan del Sur, Lanao del Norte, Misamis Oriental, Zamboanga City), Negros (Prov.: Negros Occidental, Negros Oriental), Palawan, Panay (Prov.: Alkan, Antique), Samar, Sulu Archipelago (Jolo, Siasi, Sibutu, Tawi-Tawi), Tumindao.

GENERAL DISTRIBUTION (OTHER THAN PHILIPPINES).— Malaysia (Borneo [Sabah, Sarawak]); Indonesia (Belitung, Borneo [Kalimantan], Buton, Kalimantan, Sangihe Archipelago, Sulawesi) (see Vogel, et al. [2007:23, 31] for details).

REMARKS.— Given the variation observed among samples of this species studied by Vogel et al. (2007), they concluded, “We refrain from giving a more detailed description here, as the variation among this species or complex of species will be discussed in the next and forthcoming paper of the series. A splitting into several taxa seems to be likely.” ( Vogel et al., 2007:23). As noted by Gaulke (1994:141), these seemingly non-aggressive but dangerously venomous snakes are actually beneficial to humans because their dietary preferences include agricultural pest species, rodents, and even large rats.

CONSERVATION STATUS [IUCN].— Least Concern [2016] ver. 3.1.

General geographic distribution of snakes in the Philippine Archipelago

Azucar Basilan

Tropidonophis negrosensis Indotyphlops braminus

Ramphotyphlops olivaceus

Babuyan Claro (Babuyan Ids.) Ramphotyphlops suluensis

Boiga philippina Malayopython reticulatus

Lycodon cf. alcalai Aplopeltura boa

Lycodon bibonius Ahaetulla prasina preocularis

Laticauda colubrina Boiga cynodon

Laticauda semifasciata Calamaria gervaisii hollandi Trimeresurus (Parias) cf. flavomaculatus Calamaria lumbricoidea

Chrysopelea paradisi variabilis

Balabac Coelognathus erythrurus erythrurus Xenopeltis unicolor Cyclocorus nuchalis nuchalis

Aplopeltura boa Dendrelaphis marenae

Ahaetulla prasina prasina Dendrelaphis philippinensis

Boiga cynodon Lycodon dumerilii

Boiga dendrophila multicincta Oxyrhabdium modestum

Chrysopelea paradisi variabilis Psammodynastes pulverulentus Coelognathus philippinus Rhabdophis auriculatus myersi Dendrelaphis levitoni Rhabdophis lineatus

Dendrelaphis marenae Tropidonophis dendrophiops

Dryocalamus philippinus Tropidolaemus subannulatus

Gonyosoma oxycephalum

Oligodon notospilus Batan (Batanes Ids.)

Psammodynastes pulverulentus Indotyphlops braminus

Rhabdophis chrysargos Python reticulatus

Calliophis bilineata Ahaetulla prasina preocularis Ophiophagus hannah Coelognathus erythrurus manillensis Trimeresurus (Parias) schultzei Gonyosoma oxycephalum

Tropidolaemus subannulatus Lycodon alcalai

Lycodon muelleri

Bantayan Psammodynastes pulverulentus Acrochordus granulatus Rhabdophis spilogaster

Cerberus schneiderii Trimeresurus (Parias) mcgregori Chrysopelea paradisi variabilis

Dendrelaphis marenae Biliran

Lycodon capucinus Calamaria lumbricoidea

Laticauda colubrina Oxyrhabdium modestum

Laticauda laticaudata Pseudorabdion mcnamarae

Rhabdophis lineatus

Banton Tropidonophis dendrophiops

Chrysopelea paradisi variabilis Bohol

Barit (Babuyan Ids.) Gerrhopilus hedraeus

Coelognathus erythrurus manillensis Gerrhopilus manilae

Laticauda colubrina Indotyphlops braminus

Ramphotyphlops cumingii

Malayopython reticulatus

Cerberus schneiderii Liopeltis philippinus

Aplopeltura boa Oligodon notospilus

Ahaetulla prasina preocularis Psammodynastes pulverulentus

Boiga angulata Rhabdophis chrysargos

Boiga cynodon Sibynophis bivittatus

Calamaria lumbricoidea Calliophis bilineata

Coelognathus erythrurus erythrurus

Naja sumatrana

Dendrelaphis marenae

Dendrelaphis philippinensis

Cagayan Sulu (Sulu Archipelago)

Gonyosoma oxycephalum

Calamaria suluensis

Lycodon capucinus

Oxyrhabdium modestum Dendrelaphis marenae

Psammodynastes pulverulentus

Rhabdophis auriculatus myersi Calamian Ids. (see also Calauit)

Malayopython reticulatus

Rhabdophis lineatus

Acrochordus granulatus

Tropidonophis dendrophiops

Naja samarensis (?) Pseudorabdion oxycephalum Ophiophagus hannah

Laticauda colubrina Calauit (Calamian Archipelago)

Laticauda semifasciata Indotyphlops braminus

Tropidolaemus subannulatus Malayopython reticulatus

Acrochordus granulatus

Bongao (Sulu Archipelago) Ahaetulla prasina prasina

Indotyphlops braminus Dendrelaphis caudolineatus caudolineatus Malayopython reticulatus Dendrelaphis marenae

Xenopeltis unicolor Coelognathus philippinus

Ahaetulla prasina suluensis Liopeltis philippinus

Chrysopelea paradisi variabilis Oligodon notospilus

Coelognathus philippinus Calliophis bilineata

Dendrelaphis flavescens Naja sumatrana

Dendrelaphis marenae

Gonyosoma oxycephalum Calayan (Babuyan Ids.)

Oligodon meyerinkii Malayopython reticulatus Psammodynastes pulverulentus Boiga dendrophila divergens

Chrysopelea paradisi variabilis

Boracay (off of Panay) Coelognathus erythrurus manillensis Indotyphlops braminus Cyclocorus lineatus lineatus Malayotyphlops castanotus Dendrelaphis luzonensis

Gonyosoma oxycephalum

Bubuan (Sulu Archipelago) Lycodon cf. alcalai

Ramphotyphlops olivaceus Oxyrhabdium leporinum leporinum Ramphotyphlops suluensis Laticauda colubrina

Chrysopelea paradisi variabilis Laticauda laticaudata

Dendrelaphis flavescens Trimeresurus (Parias) cf. flavomaculatus Liopeltis tricolor

Oligodon vertebralis notospilus Camiguin (unspecified Norte or Sur)

Indotyphlops braminus

Busuanga (Calamian Archipelago) Chrysopelea paradisi variabilis Indotyphlops braminus Dendrelaphis marenae

Ahaetulla prasina prasina Psammodynastes pulverulentus Coelognathus philippinus Tropidonophis dendrophiops Dendrelaphis marenae Trimeresurus (Parias) flavomaculatus

Trimeresurus (Parias) mcgregori Ptyas luzonensis

Rhabdophis spilogaster

Camiguin Norte (Babuyan Ids.) Hemibungarus mcclungi

Indotyphlops braminus Ophiophagus hannah

Malayotyphlops luzonensis Trimeresurus (Parias) flavomaculatus Malayotyphlops ruficaudus

Ahaetulla prasina preocularis Cebu

Boiga cynodon Gerrhopilus hedraeus

Calamaria gervaisii Indotyphlops braminus

Cyclocorus lineatus lineatus Malayotyphlops hypogius Dendrelaphis luzonensis Malayotyphlops luzonensis Gonyosoma oxycephalum Ramphotyphlops cumingii

Lycodon cf. alcalai Malayopython reticulatus

Lycodon bibonius

Acrochordus granulatus

Lycodon chrysoprateros

Cerberus schneiderii

Rhabdophis spilogaster

Ahaetulla prasina preocularis Trimeresurus (Parias) cf. flavomaculatus

Calamaria gervaisii iridiscens

Chrysopelea paradisi variabilis Camiguin Sur

Coelognathus erythrurus psephenourus Ahaetulla prasina preocularis

Cyclocorus lineatus alcalai

Calamaria lumbricoidea

Dendrelaphis fuliginosus Coelognathus erythrurus erythrurus

Dendrelaphis marenae

Oxyrhabdium modestum

Dendrelaphis philippinensis Tropidonophis dendrophiops

Calliophis philippinus Lycodon capucinus

Oxyrhabdium leporinum visayanum Camotes Ids. (see Pacijan, Poro) Psammodynastes pulverulentus

Pseudorabdion mcnamarae Candaraman Pseudorabdion montanum Dendrelaphis levitoni Pseudorabdion oxycephalum Dendrelaphis marenae Tropidonophis negrosensis

Hemibungarus gemianulis

Capones Ophiophagus hannah

Laticauda semifasciata Hydrophis cyanocinctus

Laticauda colubrina

Carabao Laticauda laticaudata

Calamaria gervaisii gervaisii

Dendrelaphis marenae Coron (Calamian Archipelago)

Lycodon capucinus Ahaetulla prasina prasina

Dendrelaphis philippinensis Catanduanes (?) Dryophiops rubescens Malayopython reticulatus

Cerberus schneiderii

Culion (Calamian Archipelago)

Boiga angulata

Ahaetulla prasina prasina

Boiga dendrophila latifasciata

Boiga cynodon

Calamaria gervaisii gervaisii

Coelognathus philippinus

Cyclocorus lineatus lineatus

Dendrelaphis marenae

Dendrelaphis marenae

Liopeltis philippinus

Dendrelaphis philippinensis

Hologerrhum philippinum Oligodon perkinsi

Lycodon muelleri Rhabdophis chrysargos

Oxyrhabdium modestum Sibynophis bivittatus

Calliophis bilineata Gigantes Sur (off of Panay)

Naja sumatrana Indotyphlops braminus

Hydrophis [ Chitulia ] ornatus

Cuyo Hydrophis [ Astrotia ] stokesii

Cerberus schneiderii

Lycodon capucinus Guimaras

Laticauda semifasciata Indotyphlops braminus

Acrochordus granulatus

Dalupiri (Babuyan Ids.) Coelognathus erythrurus psephenourus Indotyphlops braminus Cyclocorus lineatus alcalai Malayotyphlops luzonensis Dendrelaphis marenae

Python reticulatus Dendrelaphis philippinensis

Boiga philippina Hemibungarus gemianulis Chrysopelea paradisi variabilis

Coelognathus erythrurus manillensis Inampulugan

Dendrelaphis luzonensis Malayotyphlops castanotus

Lycodon chrysoprateros Boiga cynodon

Laticauda colubrina Coelognathus erythrurus psephenourus Trimeresurus (Parias) cf. flavomaculatus Cyclocorus lineatus alcalai

Dinagat Itbayat

Cerberus schneiderii Lycodon muelleri

Ahaetulla prasina preocularis Malayopython reticulatus

Boiga cynodon

(?) Boiga dendrophila latifasciata Ivojos (Babuyan Ids.)

Calamaria lumbricoidea Indotyphlops braminus

Calliophis salitan

Chrysopelea paradisi variabilis Jolo (Sulu Archipelago)

Gonyosoma oxycephalum Indotyphlops braminus

Lycodon dumerilii Malayopython reticulatus Oxyrhabdium modestum Xenopeltis unicolor

Psammodynastes pulverulentus Cerberus schneiderii

(?) Pseudorabdion oxycephalum Ahaetulla prasina preocularis Rhabdophis lineatus Calamaria joloensis

Stegonotus muelleri Chrysopelea paradisi variabilis Tropidonophis dendrophiops Coelognathus erythrurus erythrurus Naja samarensis Dendrelaphis marenae

Ophiophagus hannah Oligodon meyerinkii

(?) Tropidolaemus philippensis Psammodynastes pulverulentus

(?) Tropidolaemus subannulatus Calliophis suluensis

Ophiophagus hannah

Dumaran Hydrophis [ Pelamis ] platurus Sibynophis bivittatus Laticauda colubrina

Laticauda laticaudata

Gato Trimeresurus (Parias) flavomaculatus Laticauda laticaudata Tropidolaemus subannulatus Laticauda semifasciata

Kalotkot

Gigantes (off of Panay) Chrysopelea paradisi variabilis Hydrophis [ Chitulia ] lamberti Dendrelaphis marenae

Hydrophis [ Pelamis ] platurus Dendrelaphis philippinensis

Leyte Boiga angulata

Malayopython reticulatus Boiga cynodon

Aplopeltura boa Boiga dendrophila divergens

Ahaetulla prasina preocularis Boiga drapiezii ssp.

Boiga angulata Boiga philippina

Boiga cynodon Calamaria bitorques

Boiga dendrophila latifasciata Calamaria gervaisii gervaisii

Calamaria lumbricoidea Chrysopelea paradisi variabilis Chrysopelea paradisi variabilis Coelognathus erythrurus manillensis Coelognathus erythrurus erythrurus Cyclocorus lineatus lineatus

Cyclocorus nuchalis taylori Dendrelaphis luzonensis

Dendrelaphis marenae Dendrelaphis marenae

Dendrelaphis philippinensis Dendrelaphis philippinensis

Gonyosoma oxycephalum Dryophiops philippina

Lycodon capucinus Gonyosoma oxycephalum

Lycodon dumerilii Hologerrhum philippinum

Oxyrhabdium modestum Lycodon capucinus

Psammodynastes pulverulentus Lycodon muelleri

Ptyas luzonensis Lycodon solivagus

Rhabdophis auriculatus auriculatus (?) Lycodon tessellatus

Myersophis alpestris

Rhabdophis lineatus

Oligodon ancorus

Stegonotus muelleri

Oligodon modestus

Tropidonophis dendrophiops

Oxyrhabdium leporinum leporinum

Naja samarensis

Psammodynastes pulverulentus Ophiophagus hannah

Pseudorabdion mcnamarae

Trimeresurus (Parias) flavomaculatus (?) Pseudorabdion oxycephalum

(?) Tropidolaemus philippensis Ptyas luzonensis

(?) Tropidolaemus subannulatus Rhabdophis auriculatus auriculatus

Rhabdophis barbouri

Lubang Rhabdophis spilogaster

Malayopython reticulatus Hemibungarus calligaster

Cyclocorus lineatus lineatus Hemibungarus mcclungi

Gonyosoma oxycephalum Naja philippinensis

Rhabdophis spilogaster Ophiophagus hannah

Hydrophis atriceps

Luzon Hydrophis [ Lapemis ] curtus

Gerrhopilus manilae Hydrophis cyanocinctus

Acutotyphlops banaorum

Hydrophis [ Chitulia ] lamberti Indotyphlops braminus

Hydrophis [ Chitulia ] ornatus Malayotyphlops andyi

Malayotyphlops collaris Hydrophis [ Pelamis ] platurus Malayotyphlops denrorum Hydrophis [Leioselama] semperi Malayotyphlops luzonensis Microcephalophis gracilis Malayotyphlops ruber Laticauda colubrina

Malayotyphlops ruficaudus Laticauda laticaudata

Malayopython reticulatus Laticauda semifasciata

Acrochordus granulatus Trimeresurus (Parias) flavomaculatus Cerberus microlepis Tropidolaemus subannulatus

Cerberus schneiderii

(?) Gerarda prevostiana Mabag (Babuyan Ids.)

Ahaetulla prasina preocularis Laticauda colubrina

Marinduque Malayopython reticulatus Indotyphlops braminus Aplopeltura boa Malayopython reticulatus Cerberus schneiderii Gerrhopilus hedraeus (?) Fordonia leucobalia Malayotyphlops luzonensis Ahaetulla prasina preocularis Malayotyphlops ruficaudus Boiga angulata Ramphotyphlops cumingii Boiga cynodon Ahaetulla prasina preocularis Boiga dendrophila latifasciata Cyclocorus lineatus lineatus Boiga drapiezii ssp. Dendrelaphis luzonensis Calamaria gervaisii hollandi Dendrelaphis marenae Calamaria lumbricoidea Dryophiops philippina Calamaria virgulata Gonyosoma oxycephalum Chrysopelea paradisi variabilis Hologerrhum philippinum Coelognathus erythrurus erythrurus Lycodon muelleri Cyclocorus nuchalis nuchalis Oxyrhabdium leporinum leporinum Cyclocorus nuchalis taylori Naja philippinensis Dendrelaphis marenae Dendrelaphis philippinensis

Maripipi (?) Dryophiops philippina Oxyrhabdium modestum Gonyosoma oxycephalum Lycodon capucinus

Marongas Lycodon dumerilii Chrysopelea paradisi variabilis Oligodon maculatus Oligodon modestus

Masbate (?) Oligodon notospilus (doubtful) Indotyphlops braminus Opisthotropis alcalai Malayotyphlops luzonensis Oxyrhabdium modestum Ramphotyphlops cumingii Psammodynastes pulverulentus Malayopython reticulatus Pseudorabdion ater Acrochordus granulatus (?) Pseudorabdion oxycephalum Cerberus schneiderii Pseudorabdion taylori Ahaetulla prasina preocularis Rhabdophis auriculatus auriculatus Calamaria gervaisii iridescens (eastern) Chrysopelea paradisi variabilis Rhabdophis auriculatus myersi Coelognathus erythrurus psephenourus (western) Dendrelaphis fuliginosus Rhabdophis lineatus (?) Dendrelaphis luzonensis Stegonotus muelleri Dendrelaphis marenae Tropidonophis dendrophiops Lycodon capucinus Calliophis philippina Pseudorabdion mcnamarae Naja samarensis Tropidonophis negrosensis Ophiophagus hannah Hemibungarus gemianulis Hydrophis atriceps Naja philippinensis Hydrophis [ Lapemis ] curtus Hydrophis cyanocinctus

Medis Hydrophis [ Pelamis ] platurus Chrysopelea paradisi variabilis Laticauda colubrina Laticauda laticaudata

Mindanao Trimeresurus (Parias) flavomaculatus Gerrhopilus hedraeus (?) Tropidolaemus hombronii Indotyphlops braminus (?) Tropidolaemus philippensis (?) Malayotyphlops luzonensis (?) Tropidolaemus subannulatus Ramphotyphlops cumingii

Mindoro Pseudorabdion montanum

Gerrhopilus hedraeus Pseudorabdion oxycephalum

Indotyphlops braminus Ptyas luzonensis

Malayotyphlops ruber Tropidonophis negrosensis

Malayopython reticulatus Hemibungarus gemianulis

Ahaetulla prasina preocularis Ophiophagus hannah

Calamaria gervaisii gervaisii Hydrophis [ Lapemis ] curtus

Chrysopelea paradisi variabilis Laticauda colubrina

Coelognathus erythrurus manillensis Laticauda semifasciata

Cyclocorus lineatus lineatus Trimeresurus (Parias) flavomaculatus Dendrelaphis fuliginosus Tropidolaemus subannulatus Dendrelaphis marenae

Dryophiops philippina Paan de Azucar

Lycodon capucinus Boiga cynodon

Lycodon muelleri Tropidonophis negrosensis

Oligodon ancorus

Oxyrhabdium leporinum leporinum Pacijan (Camotes Ids.)

Tropidonophis negrosensis Gerrhopilus hedraeus

Hemibungarus calligaster Malayotyphlops luzonensis

Naja philippinensis Malayotyphlops ruber

Ophiophagus hannah Coelognathus erythrurus erythrurus Laticauda laticaudata

Trimeresurus (Parias) flavomaculatus Palawan

Indotyphlops braminus

Negros Malayopython reticulatus

Gerrhopilus hedraeus Xenopeltis unicolor

Indotyphlops braminus Acrochordus granulatus

Malayotyphlops canlaonensis Aplopeltura boa

Malayotyphlops castanotus Cerberus schneiderii

Malayotyphlops luzonensis Gerarda prevostiana

Malayotyphlops ruficaudus Ahaetulla prasina prasina Ramphotyphlops cumingii Boiga cynodon

Malayopython reticulatus Boiga dendrophila multicincta Acrochordus granulatus Boiga schultzei

Cerberus schneiderii Calamaria palavanensis

Ahaetulla prasina preocularis Calamaria virgulata

Boiga angulata Chrysopelea paradisi variabilis

Boiga cynodon Coelognathus philippinus

Calamaria gervaisii iridiscens Dendrelaphis levitoni

Chrysopelea paradisi variabilis Dendrelaphis marenae

Coelognathus erythrurus psephenourus Dryocalamus philippinus

Cyclocorus lineatus alcalai Gonyosoma oxycephalum

Dendrelaphis fuliginosus Liopeltis philippinus

Dendrelaphis marenae Liopeltis tricolor

Dryophiops philippina Lycodon sealei

Gonyosoma oxycephalum Oligodon notospilus

Lycodon capucinus Opisthotropis typica

Oligodon modestus Psammodynastes pulverulentus Oxyrhabdium leporinum visayanum Ptyas carinata

(?) Oxyrhabdium modestum Rhabdophis chrysargos

Psammodynastes pulverulentus Sibynophis bivittatus

Pseudorabdion mcnamarae Calliophis bilineata

Naja sumatrana Polillo

Ophiophagus hannah Indotyphlops braminus

Laticauda semifasciata Ramphotyphlops cumingii Trimeresurus (Parias) flavomaculatus Malayopython reticulatus Trimeresurus (Parias) schultzei Cerberus schneiderii

Tropidolaemus subannulatus Ahaetulla prasina preocularis

Boiga angulata

Panay Boiga cynodon

Indotyphlops braminus Boiga dendrophila divergens Malayotyphlops castanotus Calamaria gervaisii polillensis

(?) Malayotyphlops hypogius Chrysopelea paradisi variabilis Malayopython reticulatus

Coelognathus erythrurus manillensis Acrochordus granulatus Cyclocorus lineatus lineatus Cerberus schneiderii Dendrelaphis marenae

Ahaetulla prasina preocularis Dendrelaphis philippinensis

Boiga angulata Hologerrhum philippinum

Boiga cynodon Lycodon muelleri

Boiga dendrophila levitoni Psammodynastes pulverulentus Boiga drapiezii ssp. Ptyas luzonensis

Calamaria bitorques Rhabdophis spilogaster

Calamaria gervaisii iridescens Hemibungarus mcclungi

Chrysopelea paradisi variabilis Ophiophagus hannah

Coelognathus erythrurus psephenourus Trimeresurus (Parias) flavomaculatus Cyclocorus lineatus alcalai

Dendrelaphis fuliginosus Poro (Camotes Ids.)

Dendrelaphis marenae Malayotyphlops luzonensis Dryophiops philippina Malayotyphlops ruber

Gonyosoma oxycephalum Coelognathus erythrurus erythrurus Hologerrhum dermali

Lycodon capucinus Romblon

Lycodon fausti Cerberus schneiderii

Oligodon modestus Boiga cynodon

Oxyrhabdium modestum Chrysopelea paradisi variabilis Psammodynastes pulverulentus Dryophiops philippina Pseudorabdion mcnamarae Lycodon capucinus

Pseudorabdion talonuran Ophiophagus hannah

Ptyas luzonensis

Laticauda colubrina

Tropidonophis negrosensis

Hemibungarus gemianulis

Sabtang (Batanes Ids.)

Ophiophagus hannah

Ahaetulla prasina preocularis Laticauda colubrina

Gonyosoma oxycephalum Trimeresurus (Parias) flavomaculatus

Lycodon alcalai

Tropidolaemus subannulatus

Psammodynastes pulverulentus

Panubolon (see also Guimaras)

Samar

Indotyphlops braminus

Indotyphlops braminus

Malayotyphlops ruber

Papahag [also as Papahang] (Sulu Archipelago)

Oligodon meyerinkii Ramphotyphlops marxi

Ramphotyphlops olivaceus

Malayopython reticulatus

Aplopeltura boa Coelognathus erythrurus erythrurus Ahaetulla prasina preocularis Calliophis suluensis

Boiga dendrophila latifasciata Tropidolaemus subannulatus

Chrysopelea paradisi variabilis

Coelognathus erythrurus erythrurus Sibay (off of Panay)

Indotyphlops braminus

Cyclocorus nuchalis taylori

Dendrelaphis marenae

Sibutu (Sulu Archipelago)

Dendrelaphis philippinensis

Ramphotyphlops olivaceus

Dryophiops philippina (?)

Ramphotyphlops suluensis

Lycodon capucinus

Malayopython reticulatus

Lycodon dumerilii

Lycodon ferroni Ahaetulla prasina preocularis Oxyrhabdium modestum Ahaetulla prasina suluensis Psammodynastes pulverulentus Boiga cynodon

Rhabdophis auriculatus auriculatus Chrysopelea paradisi paradisi Rhabdophis lineatus Coelognathus philippinus

Stegonotus muelleri Dendrelaphis ? ( caudolineatus ) Tropidonophis dendrophiops Oligodon meyerinkii

Calliophis philippina Hydrophis [ Pelamis ] platurus

Naja samarensis Tropidolaemus subannulatus

Hydrophis atriceps

Laticauda laticaudata Sibuyan

Trimeresurus (Parias) flavomaculatus Indotyphlops braminus

(?) Tropidolaemus philippensis Malayotyphlops ruficaudus

(?) Tropidolaemus subannulatus Ramphotyphlops cumingii

Chrysopelea paradisi variabilis

Sanga Sanga (Sulu Archipelago) Dryophiops philippina

Xenopeltis unicolor Gonyosoma oxycephalum

Ahaetulla prasina suluensis Pseudorabdion mcnamarae

Chrysopelea paradisi variabilis

Coelognathus philippinus Sicogon

Dendrelaphis flavescens Ramphotyphlops cumingii

Tropidonophis negrosensis

Semirara (off of Panay)

Indotyphlops braminus Siquijor

Malayotyphlops luzonensis Malayotyphlops luzonensis

Lycodon capucinus Boiga cynodon

Chrysopelea paradisi variabilis

Siargao Dendrelaphis marenae

Malayopython reticulatus Dryophiops philippina

Boiga dendrophila latifasciata Tropidonophis dendrophiops

Chrysopelea paradisi variabilis Trimeresurus (Parias) flavomaculatus

(?) Cyclocorus nuchalis taylori

Dendrelaphis marenae Sicogon

Lycodon dumerilii Ramphotyphlops cumingii Psammodynastes pulverulentus Tropidonophis negrosensis

Siasi (Sulu Archipelago) Sitanki (Sulu Archipelago)

Malayopython reticulatus Laticauda colubrina

Ahaetulla prasina suluensis

Sulu Archipelago (unspecified ids.) (?) Hydrophis [ Leioselasma ] spiralis Calamaria virgulata

Hydrophis atriceps Tawitawi (Sulu Archipelago)

Malayopython reticulatus

Sulu (= Jolo, Sulu Archipelago) Ahaetulla prasina suluensis

Laticauda laticaudata Boiga cynodon

Boiga drapiezii ssp.

Laticauda semifasciata

Chrysopelea paradisi variabilis

Surigao Coelognathus philippinus

Dendrelaphis marenae Dendrelaphis flavescens

Hydrophis [ Pelamis ] platurus Dendrelaphis marenae

Oligodon meyerinkii

Tablas Sibynophis geminatus geminatus Indotyphlops braminus Tropidolaemus subannulatus

Gerrhopilus hedraeus

Malayotyphlops ruber Ticao

Malayotyphlops ruficaudus (?) Dendrelaphis luzonensis

Ahaetulla prasina preocularis

Tumindao

Boiga cynodon

Tropidolaemus subannulatus

Calamaria gervaisii gervaisii

Chrysopelea paradisi variabilis

Visayan Sea area

Coelognathus erythrurus psephenourus

Hydrophis atriceps

Cyclocorus lineatus alcalai

Hydrophis cyanocinctus

Dendrelaphis marenae

Hydrophis [ Lapemis ] curtus

Lycodon capucinus

Laticauda semifasciata

Oligodon modestus

Pseudorabdion mcnamarae

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BROWN, R.M., C.D. SILER, C.H. OLIVEROS, J.A. ESSELSTYN, A.C. DIESMOS, P.A. HOSNER, C.W. LINKEM, A.J. BARLEY, J.R. OAKS, M.B. SANGUILA, L.J. WELTON, D.S. BLACKBURN, R.G. MOYLE, A.T. PETERSON, AND A.C. ALCALA. 2013. Evolutionary processes of diversification in a model island archipelago. Annual Review of Ecology, Evolution, and Systematics 44: 411–435.

BROWN, RAFE M., UTPAL SMART, ALAN E. LEVITON, AND ERIC N. SMITH. 2018. A new species of Long-gland- ed Coralsnake of the Genus Calliophis (Squamata: Elapidae) from Dinagat Island, with notes on the biogeography and species diversity of Philippine Calliophis and Hemibungarus. Herpetologica 74(1 [March; online 06Feb.2018]): 89–104, 8 figs., 3 tables. (online: <http://hljournals.org/doi/ full/ 10.1655/Herpetologica-D-17-00008?code=herl-site>) (Accessed 06 Feb. 2018)

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BROWN, WALTER C., AND ANGEL C. ALCALA. 1970. The zoogeography of the herpetofauna of the Philippine Islands, a fringing archipelago. Proceedings of the California Academy of Sciences, ser. 4, 38: 105–130.

BROWN, WALTER C., AND ALAN E. LEVITON. 1961. Discovery of the snake genus Opisthotropis in the Philippine Islands, with description of a new species. Occasional Papers of the Natural History Museum of Stanford University (8): 1–5.

BURGER, WILLIAM LESLIE. 1971. Genera of Pitvipers. Ph.D. University of Kansas. 186 pp., 9 figs.

CANTOR, THEODORE E. 1836. Sketch of an undescribed hooded serpent (Hamadryas hannah). Asiatic Researches 19: 87–93, 3 pls.

[NB: Volume 19 of Asiatic Researches was published in 1836, and plates 1 through 9, associated with the pre-Cantor articles, bear imprint volume number XIX, yet the plates associated with Cantor’s paper are imprinted “Vol. XX.” It is possible that Cantor’s three plates were not printed with his article that appeared in vol. 19, and which likely appeared in the first half of 1836, but were included in vol. 20, which was opened later that same year. Asiatic Researches was first issued in 1788 as “ Asiatic Researches; or, Transactions of the Society, instituted in Bengal, for enquiring into the History, the Antiquities, the Arts and Sciences, and Literature of Asia. Volume the First.” and printed in Calcutta by Manuel Cantopher. The first volume was reprinted in London and bears a date of 1798. Volume 19, part 1 was printed in Calcutta in 1836 by the Bengal Military Orphan Press, Calcutta under the name “Asiatic Researches”.]

CANTOR, THEODORE E. 1838. “ A notice of the Hamadryas, a genus of Hooded Serpents with poisonous fangs and maxillary teeth.” Proceedings of the Zoological Society of London 1838(part 4): 72–76.

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COPE, EDWARD D. 1861. Catalogue of the Colubridae in the museum of the Academy of Natural Sciences of Philadelphia. Part 3. Proceedings of the Academy of Natural Sciences of Philadelphia 12: 553–566.

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DE LANG, RUND, AND GERNOT VOGEL. 2005. The Snakes of Sulawesi: A Field Guide to the Land Snakes of Sulawesi with Identification Keys. Chimaira Buchhandelsgesellschaft, Frankfurt am Main, Germany. 321 pp., 189 figs.

DE LEON, WALFRIDO. 1946. Cobra venom — its possibilities in Philippine medicine. Journal of the Philippine Medical Association 22 (1): 1–6.

DE LEON, WALFRIDO, AND ENRIQUE SALAFRANCA. 1957 [1956]. Cobra-anti-venom serum production at the Alabang Serum and Vaccine Laboratories. Philippine Journal of Science 85 (4): 477–487. NB: Cover dated December 1956 but published 5 August 1957.]

DERANIYAGALA, PAULUS E. P. 1960. The taxonomy of the cobras of Southeastern Asia. Spolia Zeylanica 29: 41–63.

DE ROOIJ, NELLY (Petronella Johanna). 1917. The Reptiles of the Indo-Australian Archipelago. II. Ophidia. E. J. Brill, Leiden, Netherlands. xiv + 334 pp., 117 figs.

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DICKERSON, ROY. 1928 b. Mammals of the Philippines. Pages 273–280 in R. Dickerson, ed., Distribution of Life in the Philippines. Bureau of Science, Manila, Philippines.

DICKERSON, ROY. 1928 c. Tertiary and Quaternary palaeography of the Philippines. Pages 76–96 in R. Dickerson, ed., Distribution of Life in the Philippines. Bureau of Science, Manila, Philippines.

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DIESMOS, A.C., A.C. ALCALA, C.D. SILER, AND R.M. BROWN. 2014. Status and Conservation of Philippine Amphibians. Pages 310–336 in H. Heatwole and I. Das, eds., Conservation Biology of Amphibians of Asia. Status of Conservation and Decline of Amphibians: Eastern Hemisphere. Natural History Publications (Borneo), Kota Kinabalu, Malaysia.

DOLOROSA, ROGER G. 2014. Notes on Mangrove Snake, Boiga dendrophila multicincta (Boulenger, 1896) in Iwahig River, Puerto Princesa City. The Palawan Scientist 6: 39–41, 1 fig.

DORIA, GIULIANO, AND MASSIMO PETRI. 2010. Pseudorabdion in the Museum of Genova with description of two new species from Sumatra and a revised key to the genus. Annali del Museo Civico di Storia Naturale di Genova 102: 187–201.

DOWLING, HERNDON GLENN. 1951 a. A proposed method of expressing scale reductions in snakes. Copeia 1951: 131–134.

DOWLING, HERNDON G. 1951 b. A proposed standard system of counting ventrals in snakes. British Journal of Herpetology 1: 97–99.

DOWLING, HERNDON G. 1958. A taxonomic study of the ratsnakes. VI. Validation of the genera Gonyosoma Wagler and Elaphe Fitzinger. Copeia 1958: 29–40.

DOWLING, HERNDON G. 1959. Classification of the Serpents: A critical review. Copeia 1959: 38–52.

DOWLING, HERNDON G, AND JAY M SAVAGE. 1960. A guide to the snake hemipenes: A survey of basic structure and systematic characteristics. Zoologica 45: 17–28.

DUELLMAN, WILLIAM E. 1958. A monographic study of the colubrid snake genus Leptodeira. Bulletin of the American Museum of Natural History 114: 1–152.

DUMÉRIL, AUGUST HENRI ANDRÉ. 1853. Prodrome de la classification des reptiles ofidiens. Mémoires de l’Académie des Sciences de l’Institut de France 23: 399–536, 2 pls.

DUMÉRIL, AUGUST MARIE CONSTANT, GABRIEL BIBRON, AND AUGUST HENRI ANDRÉ DUMÉRIL. 1854 a. Erpétologie générale ou histoire naturelle complète des reptiles. Tome Septième-Primière Partie. Librairie Encyclopédique de Roret, Paris, France. xvi + 780 pp.

DUMÉRIL, AUGUST MARIE CONSTANT, GABRIEL BIBRON, AND AUGUST HENRI ANDRÉ DUMÉRIL. 1854 b. Erpétologie générale ou histoire naturelle complète des reptiles. Tome Septième-Deuxième Partie. Librairie Encyclopédique de Roret, Paris, France. xii + 781– 1536 pp.

DUNN, EMMETT REID. 1928. A tentative key and arrangement of the American genera of Colubridae. Bulletin of the Antivenin Institute of America 2: 18–24.

DUNN, EMMETT REID. 1951. The status of the snake genera Dipsas and Sibon, a problem for “Quantum evolution ”. Evolution 5: 355–358.

DUNN, EMMETT REID, AND MERLE TAYLOR DUNN. 1940. Generic names proposed in herpetology by E.D. Cope. Copeia 1940: 69–76.

DUNSON, WILLIAM A. 1978. Role of the skin in sodium and water exchange of aquatic snakes placed in seawater. American Journal of Physiology 235 (3):R 151–R159.

DUNSON, WILLIAM A., AND MARGARET K. DUNSON. 1979. A possible new salt gland in a marine homalopsid snake (Cerberus rhynchops). Copeia 1979 (4): 661–672.

DUNSON, WILLIAM A., AND SHERMAN A. MINTON JR. 1978. Diversity, distribution, and ecology of Philippine marine snakes (Reptilia, Serpentes). Journal of Herpetology 12 (3): 281–286.

EVANS, GEORGE HENRY. 1903. The king-cobra, or hamadryad- Naia bungarus (Boulenger), Ophiophagus elaps (Günther). Journal of the Bombay Natural History Society 14: 409–418.

FERNER, JOHN W., RAFE M. BROWN, ROGELIO V. SISON, AND ROBERT S. KENNEDY. 2001. The amphibians and reptiles of Panay Island, Philippines. Asiatic Herpetological Research 9: 34–70.

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FISCHER, JOHANN GUSTAV. 1882. Herpetologische Bemerkungen. I. Bemerkungen über einzelne Stücke der Schlangen-Sammlung des Kön. Zoologischen Museums in Dresden. Archiv für Naturgeschichte 48 (3): 282–286.

FISCHER, JOHANN GUSTAV. 1885. Ichthyologische und Herpetologische Bemerküngen. IV: Ueber eine Kollection von Amphibien und Reptilien von Mindanao. Jahrbuch Hamburg Wissenschaft Anstalt 2: 80–81.

FITZINGER, LEOPOLDO JOSEPH FRANZ JOHANN. 1826. Neue Classification der Reptilien nach ihren naturlichen Verwandtschaften nebst eine verwandtschaft–Tafel und einem verzeichnisse der Reptilien–Sammlung der K. K. zoologisches Museum zu Wien. J.G. Heubner, Wien [Vienna], Austria. viii + 66 pp., 1 foldout pl.

FITZINGER, LEOPOLDO JOSEPH FRANZ JOHANN. 1843. Systema Reptilium. Braunmüller et Seidel, Vindobonae (Wien [Vienna]), Austria. 106 + iv + (3) pp.

FORCART, LOTHAR. 1954. Die Taxonomie von Lycodon florensis Rooij und Stegonotus sutteri Forcart (Ophidia, Colubridae). Verhandlungen der naturforschenden Gesellschaft in Basel 65 (1): 7–8.

GARMAN, SAMUEL W. 1881. New or little-known reptiles and fishes in the museum collection. Bulletin of the Museum of Comparative Zoology, Harvard University 8 (3): 85–93.

GAULKE, MAREN. 1986. Zum Beutefangverhalten der Schumuckbaumschlange Chrysopelea paradisi Boie, 1829 (Serpentes: Colubridae). Salamandra 22 (2/ 3): 211–212.

GAULKE, MAREN. 1987. Herpetologische Beobachtungen auf der Insel Calauit (Philippinen). Die Aquarien und Terrarien Zeitschrift 7 (40): 319–322.

GAULKE, MAREN. 1993 a. First record of the polyodont snake Sibynophis geminatus geminatus (Boie, 1826) from the Philippines, with a discussion of Sibynophis bivittatus (Boulenger, 1894). Herpetological Journal 3 (4): 151–152, tables 1–2.

GAULKE, MAREN. 1993 b. Zur Taxonomie und Biologie von Oligodon meyerinkii (Steindachner, 1891). Sauria 15 (3): 3–6, 3 figs.

GAULKE, MAREN. 1994 a. Eine neue Unterart des Malaysischen Baumschnüfflers, Ahaetulla prasina suluensis n. subsp. Senckenbergiana Biologica 73 (1–2 [15 Apr.]): 45–47, 1 table.

GAULKE, MAREN. 1994 b. Contribution to the snake fauna of the Sulu Archipelago, with the description of a new subspecies of Dendrelaphis caudolineatus (Gray, 1834). Herpetological Journal 4 (4): 136–144.

GAULKE, MAREN. 1994 c. Notes on the herpetofauna of Panaon and Samar, east Visayans, Philippines. Hamadryad 19: 1–10.

GAULKE, MAREN. 1995 a. Der Sulu Archipel - Besiedlungsgeschicte, Geologie und Herpetofauna. Natur und Museum 127 (7): 217–226.

GAULKE, MAREN. 1995 b. Observations on arboreality in a Philippine blind snake. Asiatic Herpetological Research 6: 45–48.

GAULKE, MAREN. 1996. Die Herpetofauna von Sibutu Island (Philippinen), unter Berücksichtigung zoogeographischer und ökologischer Aspekte. Senckenbergiana Biologica 75 (1/ 2): 45–56, 7 figs., 2 tables.

GAULKE, MAREN. 1999. Die Herpetofauna von Calauit Island (Calamianes-Inseln, Provinz Palawan, Philippinen) (Amphibia et Reptilia). Faunistische Abhandlungen, Staatliches Museum für Tierkunde Dresden 21 (19 [15 July]): 273–282, 1 fig. [map].

GAULKE, MAREN. 2001. Die Herpetofauna von Sibaliw (Panay), einem der letzten Tieflandregenwaldgebiete der West-Visayas, Philippinen — Teil II: Schlangen. Herpetofauna (Weinstadt) 23 (131): 23–34.

GAULKE, MAREN. 2002. A new species of Lycodon from Panay Island, Philippines (Reptilia, Serpentes, Colubridae). Spixiana 25 (1): 85–92.

GAULKE, MAREN. 2011. The Herpetofauna of Panay Island, Philippines. Chimaira Buchhandelsgesellschaft, Frankfurt am Main, Germany. 390 pp., 258 figs.

GAULKE, MAREN, AND ALEXANDER V. ALTENBACH. 1994. Contribution to the knowledge of the snake fauna of Masbate (Philippines) (Squamata: Serpentes). Herpetozoa 7 (1/ 2): 63–66.

GAULKE, MAREN, ARNOLD DEMEGILLO, JOCHEN REITER, AND BENJAMIN TACUD. 2003. Additions to the herpetofauna of Panay Island, Philippines. Salamandra 39 (2): 111–122.

GAULKE, MAREN, ARNOLD DEMEGILLO, AND GERNOT VOGEL. 2004. Eine neue Unterart der Mangroven-Nachtbaumnatter von den Philippinen [A new subspecies of the Mangrove snake from the Philippines]. Herpetofauna (Weinstadt) 25 (143): 5–16.

GAULKE, MAREN, AND GERSOM OPERIANO. 2006. Oxyrhabdium leporinum visayanum *, Gebänderte Philippinische Wühlschlange [Oxyrhabdium leporinum visayanum *, Banded Philippine Burrowing Snake]. Sauria 28 (3): 51–52.

GAULKE, MAREN, AND GERNOT VOGEL. 2005. Verbreitungsnachweis und Längenrekord Calamaria bitorques Peters, 1872. Sauria 27 (1): 19–23, 6 figs.

GIRARD, CHARLES FRÉDÉRIC. 1857. Descriptions of some new reptiles, collected by the U.S. Exploring Expedition under the command of Capt. Charles Wilkes, U.S.N. Third Part.— Including the species of Ophidians, exotic to North America. Proceedings of the Academy of Natural Sciences of Philadelphia 1857(August): 181–182.

GIRARD, CHARLES FRÉDÉRIC. 1858. Herpetology. In: United States Exploring Expedition, during the years 1838, 1839, 1840, 1841, 1842, under the command of Charles Wilkes, U.S.N. Vol. XX. C. Sherman & Son, Philadelphia, Pennsylvania, USA. 17 + 496 pp.

GISTEL, JOHANNES VON NEPOMUK FRANZ XAVER. 1848. Naturgeschichte des Thierreichs für höhere Schulen. Scheitlin & Krais, Stuttgart, Germany. xvi + 216 pp.

[NB: No specific Philippine listings. Adler notation on his photocopy of this paper: “Acad. Nat. Sci. Phila. has copy of ed. 1 (1848).” According to Adler, the 1851 paper (of which he has a photocopy) is identical to the 1848 publ. Adler also notes, “see Dubois 1987 Alytes 6: 45–46 for review of Gistel’s names.”; 617 figs., 32 pls.]

GOLAY, PHILIPPE. 1985. Checklist and Keys to the Terrestrial Proteroglyphs of the World (Serpentes: Elapidae-Hydrophiidae). Fondation culturelle Elapsoidea, Genève, Switzerland. ix + 91 pp.

GOLAY, PHILIPPE, HOBART M. SMITH, DONALD G. BROADLEY, JAMES R. DIXON, COLIN MCCARTHY, JEAN-CLAUDE RAGE, BEAT SCHÄTTI, AND MICHIHISA TORIBA. 1993. Endoglyphs and Other Major Venomous Snakes of the World: A Checklist. Azemiops S.A., Herpetological Data Center, Aïre-Geneva, Switzerland. xv + 478 pp.

GORMAN, GEORGE C., PAUL LICHT, AND FRED MCCOLLUM. 1981. Annual reproductive pattern in three species of marine snakes from the central Phillippines [sic]. Journal of Herpetology 15 (3): 335–354.

GRAY, JOHN EDWARD. 1842. Synopsis of the species of rattle snakes, or family of Crotalidae. The Zoological Miscellany (2): 47–51.

GRAY, JOHN EDWARD. 1843. Description of two new species of reptiles from the collection made during the voyages of H. M.S. Sulfur. Annals and Magazine of Natural History 11 (57): 46.

GRAY, JOHN EDWARD. 1849. Catalogue of the Specimens of Snakes in the Collection of the British Museum. Trustees of the British Museum, London, UK. xv + 125 pp.

GREENE, HARRY W. 1989. Defensive behaviour and feeding biology of the Asian mock viper, Psammodynastes pulverulentus (Colubridae), a specialized predator on scincoid lizards. Chinese Herpetological Research 2 (2): 21–32.

GRESSITT, JUDSON LINSLEY. 1937. Note on a Philippine cobra. Copeia 1937 (1): 73.

GRIFFIN, LAWRENCE E. 1909 a. Two new species of snakes found in the Philippine Islands. Philippine Journal of Science 4 (1): 55–56.

GRIFFIN, LAWRENCE E. 1909 b. Poisonous snakes of the Philippine Islands. Philippine Journal of Science 4 (3): 203–204.

GRIFFIN, LAWRENCE E. 1909 c. A list of snakes found in Palawan. Philippine Journal of Science 4 (6): 595–601.

GRIFFIN, LAWRENCE E. 1910. A list of snakes from the island of Polillo, P.I., with descriptions of a new genus and two new species. Philippine Journal of Science 5 (4): 211–218.

GRIFFIN, LAWRENCE E. 1911. A check-list and key of Philippine snakes. Philippine Journal of Science 6 (5): 253–268.

GRITIS, PAUL, AND HAROLD K. VORIS. 1990. Variability and significance of parietal and ventral scales in the marine snakes of the genus Lapemis (Serpentes: Hydrophiidae), with coments on the occurrence of spiny scales in the genus. Fieldiana: Zoology (56):i-iii, 1–13.

GUICHENOT, ALPHONSE. 1853. (see Jacquinot, Honoré, and Alphonse Guichenot. 1853)

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GÜNTHER, ALBERT C.L.G. 1859. On the genus Elaps Wagler. Proceedings of the Zoological Society of London 1859: 79–89.

GÜNTHER, ALBERT C.L.G. 1860. Note on Psammophis Perroteti, D. & B. Annals and Magazine of Natural History 6: 428–429.

GÜNTHER, ALBERT C.L.G. 1862. On new species of snakes in the collection of the British Museum. Annals and Magazine of Natural History 9: 124–132.

GÜNTHER, ALBERT C.L.G. 1864. The Reptiles of British India. Robert Hardwick (for the Ray Society), London, England, UK. xxvii + 452, 26 pls.

GÜNTHER, ALBERT C.L.G. 1873. Notes on some reptiles and batrachians obtained by Dr. Adolf Bernhard Meyer in Celebes and the Philippine Islands. Proceedings of the Zoological Society of London 1873: 165–172, 2 pls.

GÜNTHER, ALBERT C.L.G. 1879. List of the mammals, reptiles, and batrachians sent by Mr. Everett from the Philippine Islands. Proceedings of the Zoological Society of London 1879 (5): 74–79.

GÜNTHER, ALBERT C.L.G. 1883. Description of two new snakes from the “Challenger” collections. Annals and Magazine of Natural History 11: 136–137.

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HAAS, C.P.J. DE. 1950. Checklist of the snakes of the Indo-Australian Archipelago (Reptiles, Ophidia). Treubia 20: 511–625.

HAAS, GEORG. 1931. Die kiefermuskulatur und die Schädelmechanik der Schlangen in Vergleichender Darstellung. Zoologische Jahrbücher. Abteilung für Anatomie und Ontogenie der Tier 53: 127–198.

HAHN, DONALD E. 1980. Liste der rezenten Amphibien und Reptilien: Anomalepididae, Leptotyphlopidae, Typhlopidae. Das Tierreich 101:i-xii+ 1–93.

HARDING, KEITH A., AND KENNETH R.G. WELCH. 1980. Venomous Snakes of the World: A Checklist. Pergamon Press, Oxford, UK. x + 188 pp.

HEANEY, L.R. 1985. Zoogeographic evidence for middle and Late Pleistocene land bridges to the Philippine Islands. Modern Quaternary Research in Southeast Asia 9: 127–143.

HEATWOLE, HAROLD, ALANA GRECHI, AND HELENE MARSH. 2017. Paleoclimatology, paleogeography, and the evolution and distribution of sea kraits (Serpentes: Elapidae: Laticauda). Herpetological Monographs 31 (1; Dec. 2017): 1–17.

HEDGES, S. BLAIR, ANGELA B. MARION, KELLY M. LIPP, JULIE MARIN, AND NICOLAS VIDAL. 2014. A taxonomic framework for typhlopid snakes from the Caribbean and other regions (Reptilia, Squamata). Caribbean Herpetology 49: 1–61, tables 1–3, Appendix.

HELFENBERGER, NOTKER. 2001. Phylogenetic relationships of Old World ratsnakes based on visceral organ topology, osteology, and allozyme variation. Russian Journal of Herpetology 8 (Suppl. 1): 1–62.

HERRE, ALBERT WILLIAM CHRISTIAN THEODORE. 1928. True fresh-water fishes of the Philippines. Pages 242–247 in R. Dickerson, ed., Distribution of Life in the Philippines. Bureau of Science, Manila, Philippines.

HERRE, ALBERT W.C.T. 1942. Notes on Philippine sea snakes. Copeia 1942 (1): 7–9.

HERRE, ALBERT W.C.T., AND D.S. RABOR. 1949. Notes on Philippine sea snakes of the genus Laticauda. Copeia 1949 (4): 282–284.

HORNSTEDT, CLAËS FREDRIK. 1787. Beskrifning på en Ny Orm från Java. Kongl. Vetenskaps Academiens nya Handlingar, Stockholm 8: 306–308.

IN DEN BOSCH, HERMAN A.J. 1985. Snakes of Sulawesi: Checklist, key and additional biogeographical remarks. Zoologische Verhandelungen, Leiden 217: 3– 50 pp.

IN DEN BOSCH, HERMAN A.J., AND IVAN INEICH. 1994. The Typhlopidae of Sulawesi (Indonesia): A review with a description of a new genus and a new species (Serpentes: Typhlopidae). Journal of Herpetology 28 (2): 206–217.

INGER, ROBERT F. 1954. Systematics and zoogeography of Philippine Amphibia. Fieldiana: Zoology 33: 181–531.

INGER, ROBERT F. 1967. A new colubrid snake of the genus Stegonotus from Borneo. Fieldiana: Zoology 51 (5): 77–83.

INGER, ROBERT F, AND PHILIP J. CLARK. 1943. Partition of the genus Coluber. Copeia 1943: 141–145.

INGER, ROBERT F., AND ALAN E. LEVITON. 1966. The taxonomic status of Bornean snakes of the genus Pseudorabdion Jan and of the nominal genus Idiopholis Mocquard. Proceedings of the California Academy of Sciences, ser. 4, 34 (4): 307–314.

INGER, ROBERT F., AND HYMEN MARX. 1962. Variation of hemipenes and cloaca in the colubrid snake Calamaria lumbricoidea. Systematic Zoology 11 (1): 32–38.

INGER, ROBERT F., AND HYMEN MARX. 1965. The systematics and evolution of the Oriental colubrid snakes of the genus Calamaria. Fieldiana: Zoology 49: 1–304, 73 figs.

INGER, ROBERT F., AND HAROLD K. VORIS. 2001. The biogeographical relations of the frogs and snakes of Sundaland. Journal of Biogeography 28: 863–891, 4 figs., 7 tables, Appendix.

ISKANDAR, D.T., AND E. COLIJN. 2002. A Checklist of Southeast Asian and New Guinean Reptiles. Part I: Serpentes. Biodiversity Conservation Project, Djakarta, Java, Indonesia.

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JAN, GEORG. 1857. Cenni sul Museo Civico di Milano ed indice sistematico dei retilli ed anfibi eposti nel medesimo. Luigi di Giacomo Pirola, Milano, Italia. 61 pp.

JAN, GEORG. 1862. Enumerazione sistematica della specie d’Ophidi del gruppo Calamaridae. Archivo Zool. Anat. Fisiol. Modena 2 (1): 1–76.

[NB: Reprinted in August, 1862 as Prodromo dell’Iconografia Generale degli Ofidi. II. Parte. V. Gruppo: Calamaridae, with the addition of an introduction by Jan, pp. iii-xii, pp. iii-x dated 20 dicembre 1861 and pp. x-xii, 1 Agosto 1862.; pls. 5–6.]

JAN, GEORG. 1863 a. Elenco sistematico degli Ofidi. Descritti e disegnati per l’Iconographia Generale. A. Lombardi, Milano, Italy. vii + 143 pp.

JAN, GEORG. 1863 b. Enumerazione sistematica degli ofidi appartenenti al gruppo Coronellidae. Archivo Zool. Anat. Fisiol. Modena 2 (2): 213–330.

[NB: Reprinted in August, 1863 as Prodromo dell’Iconografia Generale degli Ofidi. II.ª Parte. VI.º Gruppo: Coronellidae, pp. (2) + 3–120.]

JENSEN, LAURITZ A., GERALD D. SCHMIDT, AND ROBERT E. KUNTZ. 1983. A survey of cestodes from Borneo, Palawan and Taiwan, with special reference to three new species. Proceedings of the Helminthological Society of Washington 50 (1): 117–134.

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KLUSMEYER, BORIS, AND BERND FAUSTEN. 1994. Haltung und Nachzucht der Bambusotter Trimeresurus flavomaculatus halieus. Salamandra 30 (3): 174–184.

KROON, CHARLES. 1973. A second specimen of the colubrid smake Boiga philippina. Herpetologica 29 (1): 51–53.

LACÉPÈDE, BERNARDE GERMAINE ÉTIENNE. 1789. Histoire Naturelle des Quadrupèdes Ovipares et des Serpens. Hôtel de Thou, Paris, France. 1–8 + 9 –19 + [20] + 1–77 + 78–144 +1– 527 pp., 22 pls.

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LANZA, BENEDITTO. 1999. A new species of Lycodon from the Philippines, with a key to the genus (Reptilia Serpentes Colubridae). Tropical Zoology 12: 89–104, 3 figs.

[NB: Includes a key to identify all species included by Lanza in the genus Lycodon.]

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[NB: See comments by Bauer & Adler (2001) on the order in which the different printings of the Meyen reports appeared. The volume cited here, according to Bauer and Adler, though stating on the title pages of each of the parts as appearing in the “ Acta Acad. Caes. Leop. Carol. Nat. Cur. Vol. XVII. P.I.” actually appeared before it appeared in the Acta. The pages in the volume bear dual pagination numbers, e.g. “436 (186),” the first as it appears in the full volume, the second the already assigned pagination for the Acta. Thus, Wiegmann’s Amphibian section is numbered 433–435 (actually these pages are unnumbered) followed by 436 (186)-522(268). In a like manner the reptile and amphibian plates are dually numbered, LII (XIII)-LXI (XXII)., pls. 52–61.]

WIEGMANN, AREND FRÉDÉRIC AUGUST HEINRICH. 1834 a. Amphibien. Pages 433–518, Beiträge zur Zoology, gesammelt auf einer Reise um die Erde von Dr. F.J.F. Meyen. Für den Verfasser in Commission den Sander’schen Buchhandlung zu Berlin (C.W. Eichhoff), Breslau und Bonn.

[NB: Author’s name shown as “Dr. A.F.A. Wiegmann dem Jüngern” on title page [p. 433 of Meyen volume]. Pagination of Nova Acta version indicated in parentheses: 433–435 [183–185 {but not shown on pages or until p. 436}], 436[186]-518[268]; pls. in Meyen vol./ Nova Acta numbered 52/13–61/22. See also Wiegmann’s “Nachträgliche Bemerkungen” chapter, numbered in Nova Acta as 268a–d. Original description and pl. of Elaps calligaster (pl. 59, fig. 2 [pl. 20, fig. 2 in Nova Acta]., pls. 52–61).]

WIEGMANN, AREND FRÉDÉRIC AUGUST HEINRICH. 1834 b. Nachträgliche Bemerkungen zur Beschreibung der Amphibien in Meyen’s Beiträgen zur Zoologie. Pages 519–522, Beiträge zur Zoology, gesammelt auf einer Reise um die Erde von Dr. F.J.F. Meyen. Für den Verfasser in Commission den Sander’schen Buchhandlung zu Berlin (C.W. Eichhoff), Breslau und Bonn.

[NB: Annotated: “Berlin im October 1834.” See also Wiegmann’s Siebente Abhandlung. Amphibien. Pagination of Nova Acta version indicated in parentheses: 433–435[183–185 {but not shown on pages or until p. 436}], 436[186]-518[268]; pls. in Meyen vol./ Nova Acta numbered 52 / 13–61 /22. See also Wiegmann’s “Nachträgliche Bemerkungen” chapter, numbered in Nova Acta as 268a–d but here as 519[268]– 522[268].]

WIEGMANN, AREND FRÉDÉRIC AUGUST HEINRICH. 1834 c. Beiträge zur Zoologie, gesammelt auf einer Reise um die Erde, von Dr. F.J.F. Meyen, M.d.A.d.N. Siebente Abhandlung. Amphibien. Nova Acta Academiae Caesareae Leopoldino-Carolinae Germanicae Naturae Curiosorum Halle 17 (1): 185–268, 268a–d, 11 pls.

[NB: According to Bauer and Adler (2001), preprints of this paper were included in F.J.F. Meyen (1834), Reise um die Erde (q.v.) and then issued in the Nova Acta, also in 1834, though somewhat later (but not much since Meyen signed the foreword to the book [pp. iii-iv] on 15 Sept. 1834).]

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CM

Chongqing Museum

Kingdom

Animalia

Phylum

Chordata

Class

Squamata

Family

Viperidae

Genus

Trimeresurus

Loc

Trimeresurus (Parias) schultzei (Griffin, 1909)

Leviton, Alan E., Siler, Cameron D., Weinell, Jeffrey L. & Brown, Rafe M. 2018
2018
Loc

Tropidolaemus philippensis, Vogel, David, Lutz, van Rooijen, and Vidal, 2007:31

SANGUILA, M. B. & K. A. COBB & C. D. SILER & A. C. DIESMOS & A. C. ALCALA & R. M. BROWN 2016: 119
LEVITON, ALAN & M. BROWN & AND CAMERON & D. SILER 2014: 518
VOGEL, GERNOT & PATRICK DAVID, MARIO & LUTZ, JOHAN & VAN ROOIJEN, AND 2007: 31
2007
Loc

Tropidolaemus subannulatus, Vogel, David, Lutz, van Rooijen, and Vidal, 2007:23

SANGUILA, M. B. & K. A. COBB & C. D. SILER & A. C. DIESMOS & A. C. ALCALA & R. M. BROWN 2016: 109
LEVITON, ALAN & M. BROWN & AND CAMERON & D. SILER 2014: 519
GAULKE, MAREN 2011: 340
VOGEL, GERNOT & PATRICK DAVID, MARIO & LUTZ, JOHAN & VAN ROOIJEN, AND 2007: 23
2007
Loc

Tropidolaemus cf. wagleri

FERNER, JOHN & M. BROWN & ROGELIO & V. SISON & AND ROBERT & S. KENNEDY 2001: 55
2001
Loc

Tropidolaemus wagleri

MCDIARMID, ROY & A. CAMPBELL & AND T'SHAKA & A. TOURE 1999: 349
GAULKE, MAREN 1994: 141
BURGER, WILLIAM 1971: 2
1971
Loc

Trimeresurus wagleri, Leviton, 1964c:265

LEVITON, ALAN 1964: 265
1964
Loc

Trimeresurus schultzei

LEVITON, ALAN & M. BROWN & AND CAMERON & D. SILER 2014: 518
TORIBA, MICHICISA 1993: 105
LEVITON, ALAN 1964: 263
TAYLOR, EDWARD 1922: 292
1922
Loc

Trimeresurus wagleri

TORIBA, MICHICISA 1993: 108
LEVITON, ALAN 1964: 265
TAYLOR, EDWARD 1922: 296
1922
Loc

Trimeresurus wagleri wagleri

TAYLOR, EDWARD 1922: 302
1922
Loc

Trimeresurus wagleri alboviridis

TAYLOR, EDWARD 1917: 366
1917
Loc

Trimeresurus philippensis

TAYLOR, EDWARD 1922: 295
GRAY, JOHN 1842: 48
1842
Loc

Trimesurus subannualtus

GRAY, JOHN 1842: 48
1842
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