Trogellus, Paulsen, 2013

Paulsen, M. J., 2013, A new genus for the Neotropical species of Aesalus Fabricius, with descriptions of eight new species (Coleoptera: Lucanidae: Aesalinae), Insecta Mundi 2013 (325), pp. 1-25 : 18-19

Phylogeny of Trogellus and Aesalini

Taxonomic sampling and specimen vouchering

Sequences were analyzed for 47 taxa selected from a larger matrix examining the world Lucanidae (Paulsen and Hawks, in prep.), with 22 outgroup and 25 ingroup taxa ( Aesalinae ). The subfamily Aesalinae is represented by all three tribes and all nine genera. Of the ten species of Trogellus , only two could not be included in the analyses: the most recent specimens of T. neotropicalis located were the original type series from 1888; I could not amplify DNA from the holotype and only known specimen of T. catrachitus . Outgroup taxa included Geotrupidae , Bolboceratidae , and Pleocomidae that were shown to be the sister group to the remaining Scarabaeoidea in Smith et al. (2006); Trogidae and Glaresidae , which comprise the sister group to Lucanidae according to Smith et al. (2006); and exemplars of the other three lucanid subfamilies (Lampriminae, Syndesinae, and Lucaninae ). Outgroup taxa were chosen that had the most complete data for all gene regions in the study.

Sequenced specimens are identified by voucher labels and are currently housed in the MJPC, UNSM, or David C. Hawks Collection (Riverside, CA; DCHC). Holotype specimens that were sequenced are deposited at the institutions indicated in their descriptions. See Table S1 for a complete listing of included sequences and voucher locations.