Indocloeon (Hindocloeon), Kluge & Suttinun, 2020

Kluge, Nikita J. & Suttinun, Chanaporn, 2020, Review of the Oriental genus Indocloeon Müller-Liebenau 1982 (Ephemeroptera: Baetidae) with descriptions of two new species, Zootaxa 4779 (4), pp. 451-484 : 468

publication ID	https://doi.org/ 10.11646/zootaxa.4779.4.1
publication LSID	lsid:zoobank.org:pub:861A7289-DD34-4174-B6B2-783FBEC8DF36
DOI	https://doi.org/10.5281/zenodo.3853340
persistent identifier	https://treatment.plazi.org/id/03B587A2-F216-FF84-ECAB-FF500E2FFA5B
treatment provided by	Plazi
scientific name	Indocloeon (Hindocloeon)
status	subgen. nov.

Treatment

Subgenus Hindocloeon subgen. n.

Hierarchical name: Hindocloeon /g(1).

( Figs 56–118 View FIGURES 56–59 View FIGURES 60–63 View FIGURES 64–67 View FIGURES 68–70 View FIGURES 71–75 View FIGURES 76–81 View FIGURES 82–90 View FIGURES 91–95 View FIGURES 96–100 View FIGURES 101–105 View FIGURES 106–111 View FIGURES 112–114 View FIGURES 115–118 )

Type species: Indocloeon indonesiae Kluge 2012 View in CoL .

Etymology. The name proceeds from the geographical name Hindostan, which some authors have used to refer to the wide area comprising India and Indochina.

Diagnosis. Larva. (1) Labrum with 1 paired midway seta between submedian setae and submarginal arc of setae ( Fig. 60 View FIGURES 60–63 ; Kluge 2012: fig. 2). (2) Incisor of left mandible terminated with 4 denticles, 2nd denticle longest; incisor of right mandible terminated with 4 denticles, 3rd denticle longest ( Figs 64–66 View FIGURES 64–67 ; Kluge 2012: figs 16–17). (3) Left mandible with setae proximad of prostheca ( Fig. 64 View FIGURES 64–67 ; Kluge 2012: fig. 16). (4) Mola of left mandible without setae on proximal angle ( Fig. 64 View FIGURES 64–67 ) (in contrast to right mandible—Fig. 65). (5) Maxillary palp consists of 3 subequally long segments ( Figs 69–70 View FIGURES 68–70 ; Kluge 2012: fig. 5). (6) Paraglossae widened from base to midlength; dorso-median setal row consists of 4–5 setae ( Fig. 74 View FIGURES 71–75 ). (7) Labial palp without point on apico-median projection of 2nd segment ( Figs 71–73 View FIGURES 71–75 ; Kluge 2012: fig. 3); exuviae of palp retains its shape. (8) Tibiae of middle and hind legs, in contrast to tibia of fore leg, has undetermined number of stout setae along outer margin, all these setae being smaller than the outer-apical seta of the fore tibia ( Fig. 77 View FIGURES 76–81 ).

Imago and subimago. (9) Antenna: distal part of scapus with hypodermal pigmented spots on lateral and median sides; distal part of pedicellus with hypodermal pigmented spots on lateral and median sides ( Fig. 92 View FIGURES 91–95 ). (10) First abdominal segment usually with a pair of contrasting brown hypodermal lateral spots just ventrad of metathoracic postnotum ( Figs 93–95 View FIGURES 91–95 , 97, 100 View FIGURES 96–100 , 114 View FIGURES 112–114 , 115, 117 View FIGURES 115–118 ). (11) Abdominal terga without such transverse stripes as in Indocloeon s. str.

Subimaginal cuticle. (12) Postsubalar sclerite with postero-dorsal angle only slightly stretched ( Fig. 91 View FIGURES 91–95 ; Kluge 2012: fig. 26).

Genitals of male imago. (13) Styliger lacks median impression ( Fig. 101 View FIGURES 101–105 ; Kluge 2012: figs 24, 28). (14) Each gonovectis is divided into a proximal portion immovably attached to penial bridge and a movably articulated distal portion; the distal portion is narrow at base and plate-like widened distally; gonovectal muscle is partly attached to proximal portion, partly to movable distal portion of gonovectis ( Fig. 101 View FIGURES 101–105 ; Kluge 2012: fig. 24).

Egg. (15) Surface without net-like relief and without sharply outlined cavities other than micropilar area ( Figs 106–111 View FIGURES 106–111 ).

Composition and distribution. Two species are placed here for certain, being known from all stages of development: Indocloeon (Hindocloeon) indonesiae Kluge 2012 and Indocloeon (Hindocloeon) continentale sp. n. Other species are known from certain stages of development only, and are placed here presumably: Indocloeon (Hindocloeon) spathasetis Kaltenbach & Gattolliat 2017 and Indocloeon (Hindocloeon) timorense Kaltenbach & Gattolliat 2017 are known as larvae only; Indocloeon (Hindocloeon) longistylus ( Demoulin 1969) is known as male imago only; Indocloeon (Hindocloeon) sp. « Vietnam » and Indocloeon (Hindocloeon) sp. «Java» are reported here as male and female subimagines only.