Anogdus tridens Peck and Cook, 2013

Peck, Stewart B. & Cook, Joyce, 2013, A revision of the species of Anogdus LeConte of the United States and Canada (Coleoptera: Leiodidae: Leiodinae: Leiodini), Insecta Mundi 2013 (290), pp. 1-27 : 19-20

Anogdus tridens Peck and Cook , new species

( Fig. 31, 32 , 35 )

Diagnostic description. Length (pronotum + elytra) = 1.64–2.04 mm; greatest width = 1.12–1.48 mm. Head punctures moderately coarse, irregularly spaced. Antennal club slender; antennomere 7 narrower than 9 and 10; antennomere 8 narrow, not disk-like, distinctly thicker on the outer side; apical anten- nomere nearly as wide as 9 and 10. Sides of pronotum weakly rounded, posterior angles broadly obtuse. Pronotal punctures fine and distantly spaced on disk, larger and more dense laterally. Elytral strial punctures round and deep, separated by ± 1 diameter; interstrial punctures minute, distant; elytral epipleura with short setae. Metasternal anterior margin with a row of large, deep punctures, distinctly larger than the shallow lateral punctures of the metasternum. Protarsi and mesotarsi of male bearing white setae ventrally. Male mesotibia curved; mesotibial process broad at base, slender, curved toward and shorter than large tibial spine. Metafemur slender; males usually with a broad expansion near the apex of the lower margin, with a small, acute tip. Anterior margins of abdominal sternites III–VII each with a row of closely spaced, round, deep punctures of about equal size. Median lobe of aedeagus ( Fig. 31, 32 ) elongate, broad, somewhat flattened dorsoventrally, curved near base; apex narrowed and flat- tened. Parameres slender, apically expanded, nearly reaching apex of median lobe. Internal sac ( Fig. 31, 32 ) with a characteristic heavily sclerotized trilobed structure.

Note. This species was illustrated by Daffner (1988, Fig. 23 and 24 ) as Anogdus obsoletus (Melsheimer) . A. tridens resembles A. obsoletus externally, but is clearly distinguished by the heavily sclerotized three-pronged structure in the internal sac.

Type material (n=65). Holotype, male, in FSCA, with label data: “ OKLAHOMA / Latimer Co. / - IV- 1990 / Karl Stephan ” ; Paratypes, 64, as follows: same data as holotype (1, FSCA) ; same data except “ IV-1989 ” (1, FSCA) ; same data (1, TAMU) ; same data except “- V-1991 ” (1, FSCA); same data except “- IV-1991 ” (1, FSCA) ; same data except “ V-1992 ” (2, FSCA) ; same data except “ VIII-1988 ” (1, FSCA) ; same data except “ V-1989 ” (5, FSCA) ; same data (1, TAMU) ; same data except “ V-2002 / K. Stephan ” (1, TAMU) ; 3, same data except “ V-1982 ” (1, TAMU) ; same data except “ V-1986 ” (3, TAMU) ; same data except “ IV-1986 ” (4, TAMU) ; same data except “ V-1988 ” (1, TAMU) ; same data except “ VIII-1988 ” (1, TAMU) ; same data except “ 5mi. W. Red Oak / 7-V-77” (1, TAMU) ; “ ARKANSAS: Polk Co / 13mi. NW Mena / Rich Mt. , 2800’/ 1-3. VI.79, S. & J. Peck / mesic oak-hickory” (1, SBPC) ; “ ARIZONA: Cochise Co. / Southwest Research / Station VII-29/31-1987/ F. Andrews & T. Eichlin / Malaise trap in/ wooded riparian/ canyon bottom” (1, CSCA) ; “ Arizona; Cochise Co. / West Stronghold / Dragoon Mts. / July 20 1973 / K. Stephan leg.” (1, FSCA) ; “ Portal / Arizona ”, “DrLenczy/ 7.1968” (4, USNM) ; “ Florida Cn / Pima Co. / Arizona ”, “DrLenczy/ 8.1976” (1, USNM) ; “ Green Valley / Pima Co. / Arizona ”, “DrLenczy/ 6.1969” (4, USNM) ; “ INDIANA: Tippecanoe / Co., Lafayette / VI.27 1990 / N.M. Downie, UV light” (1, FMNH) ; “ ILLINOIS: Champaign Co., Mahomet / Nettie Hart Mem. Woods/ 22 May 66 uv/ M.W. Sanderson ” (1, SBPC) ; same data except “24 Aug 66 uv” (1, SBPC) ; same data except “18 July 66 uv” (1, SBPC) ; same data except “24 June 66 uv” (1, SBPC); Champaign Co, Urbana, Ill.”, “ May 31, 1938 / Eugene Ray” (1, FMNH) ; “ USA: TEXAS: Bell Co. / 1 km. W Youngsport, FIT/ Bowmer Rch., Lampasas Riv. / 30.96197°N, 97.72903°W / IV-10-21-2010, T. Robbins” (1, TAMU) ; “ USA: TEXAS: Brazos Co. / College Sta., Lick Creek / Park ( Site 1), FIT-ground/ 30.55841°N, 96.20866°W / bottomland forest/ V-22-VI-29-2011 / E.G. Riley,” (1, TAMU) ; “ USA: TX: Brazos Co. / Koppe’s Bridge , 5miSW/ College Sta. 17.IV-18. V.87/ Ripar- ian ravine FIT/ R. Anderson” (10, SBPC) ; “ USA Texas Brewster Co. / Big Bend National Park / Maple Cyn. 5200’/ 9.VII.82 G.A.P. Gibson ” (1, SBPC) ; “USA: TEXAS: Ellis Co. / 3.6 mi. NE Italy / 32.21752°N, 96.93389°W,/ IV-2-V-8-2008, FIT-ground/ Coll. E.G. Riley, et al.” (4, TAMU) ; “TEX. Palmetto St. / Pk. Gonzales Co. / X.73” (1, SBPC) .

Distribution. Widely distributed in the eastern United States from Illinois and Indiana through Ar- kansas to Arizona, Oklahoma and Texas ( Fig. 35 ).

Seasonality. Adults are mostly known from the months of May through August suggesting that the species is active during the monsoon rainy season in the summer months.

Bionomics. The dominant habitat is deciduous moist forest, ranging into wooded riparian canyon bottoms and open juniper scrub.

Etymology. The epithet “ tridens ” refers to the characteristic heavily sclerotized three-pronged structure in the internal sac of the aedeagus of this species.

Figures

Figure 35. North American distribution of Anogdus puritanus (circles), A. alachua (cross), A. texanus (squares), A. tridens (triangles).

Figures 19–32. Aedeagi, dorsal and lateral, of Anogdus species. 19, 20) A. superans. 21, 22) A. alachua. 23, 24) A. cochise. 25, 26) A. huachuca. 27, 28) A. rileyi. 29, 30) A. texanus. 31, 32) A. tridens. Scale line = 0.30 mm for superans, alachua, huachuca, texanus, tridens; 0.35 mm for cochise, rileyi.