Physopelta (Neophysopelta) gutta gutta ( Burmeister, 1834 )

Stehlík, Jaroslav L., 2013, Review and reclassification of the Old World genus Physopelta (Hemiptera: Heteroptera: Largidae), Acta Entomologica Musei Nationalis Pragae 53 (2), pp. 505-584 : 528-532

Physopelta (Neophysopelta) gutta gutta ( Burmeister, 1834)

( Figs 17–34 , 98 )

Lygaeus (Pyrrhocoris Fall., Platynotus Schill.) gutta Burmeister, 1834: 300–301 , pl. XLI: fig. 10.

SYNTYPE (s): Philippines, Manila env. (ZMHB, 1 ♂).

Pyrrhocoris gutta: BURMEISTER (1835) : 285 (new combination, diagnosis, distribution).

Physopelta bimaculata Stål, 1855:186 (description, distribution).

SYNTYPE (s): Indonesia, Java (NHRS, G. Lindberg, pers. comm.).

Synonymized by STÅL (1861: 195).

Physopelta gutta: STÅL (1861) : 195 (new synonymy, list); STÅL (1863): 391 (catalogue, distribution); STÅL (1870): 99–100 (key to species, catalogue, distribution); STÅL (1871): 665 (list of species); WALKER (1873): 18, 19 (key, catalogue, distribution); SCOTT (1874): 291 (distribution); DISTANT (1879a): 127 (distribution); DISTANT (1879b): 37 (distribution); DISTANT (1883): 417 (distribution); LETHIERRY (1888): 463 (distribution); KIRBY (1891): 105 (distribution); LETHIERRY & SEVERIN (1894): 242 (catalogue, distribution); UHLER (1897): 265 (variability, distribution); BREDDIN (1900): 161 (distribution; not distinguishing Ph. g. famelica ); BREDDIN (1901b): 139 (distribution); DISTANT (1903a): 97 (redescription, figures, distribution); DISTANT (1903b): 250 (catalogue, distribution); MAXWELL- LEFROY (1909): 325 (distribution); ESAKI (1926): 157 (distribution); TAEUBER (1927): 174–177 (redescription, variability, distribution; differences from Ph. lisae and Ph. roseni , respectivelly); BLÖTE (1931): 99–100 (distribution); SCHMIDT (1931): 46 (distribution); LIU (1981): 222, 224, pl. 28 (key, redescription, figure); AHMAD & ABBAS (1987): 137–138, 141–142 (redescription, figures, host plant, distribution); AHMAD & PERVEEN (1986): 162, 164–166 (morphology of abdomen and genitalia); AHMAD & PERVEEN (1989): 49–50 (cephalic morphology); ZAMAL & CHOPRA (1990): 4 –5 (redescription, figures, distribution); LEE & KWON (1991): 50 (distribution); TOMOKUNI (1993): 59, 197 (redescription, habitus, larva, egg, biology); SEN et al. (1998): 331, 333–334, 339–340 (checklist, key, diagnosis, distribution); HUA (2000):187 (check-list, host plants, distribution); KWON et al. (2001): 301–302 (check-list, host plants, distribution); TOMOKUNI (2006): 352 (distribution); SAHA & BAL (2007):322–324 (checklist, key, diagnosis, distribution); CHANDRA et al. (2012):70–71 (figure, distribution); CHANDRA & KUSHWAHA (2012): 253 (distribution); CHANDRA & KUSHWAHA (2013): 678 (distribution); SINGH & BANYAL (2013): 1056 (distribution).

Physopelta gutta gutta: HUSSEY (1929) : 30 –31 (catalogue, distribution); ESAKI (1952): 230 (redescription, habitus, distribution); STEHLÍK (1965a): 287 (identification of gutta and famelica , distribution); MANNA et al. (1985): 621–630 (chromosomes, figures, distribution); CHAKRABARTY et al. (1994): 34–35 (differential diagnosis, distribution); KERZHNER (2001): 246 (catalogue, distribution); CASSIS & GROSS (2002): 626–627 (catalogue, distribution – mixed with gutta famelica ); KOHNO et al. (2002): 393 (distribution); STEHLÍK & JINDRA (2003): 7 (distribution); STEHLÍK (2004): 2–3 (description of 5th larval instar, description); STEHLÍK (2005a): 144 (distribution); STEHLÍK (2007a): 117–118, 127 (habitus, variability, identification and distribution of gutta and famelica ); RÉDEI et al. (2009):8, 12–15, 48 (redescription, key, figures, distribution); KMENT & VILÍMOVÁ (2010):4, 11–12 (morphology of external scent efferent system).

Physopelta (Physopelta) gutta gutta: VOIGT (2006) : 224 (key, distribution – mixed with gutta famelica ).

Physopelta (Physopelta) gutta famelica (misidentification): VOIGT (2006): 224 (record from China).

Physopelta guttata (incorrect subsequent spelling): YANG (2003): 200–201 (morphology, figures).

Material examined. JAPAN: KYUSHU: Korasan, Chikugo, 1.viii.1955, 1 ♂, 25.ix.1955, 1 ♀, S. Miyamoto lgt. & det. (NMPC). – RYUKYUS: OKINAWA ISLAND: W slope of Katsudake Motobu Pen., 28.ix.1945, 1 ♂, F. G. Werner lgt. (AMNH); ditto, Iwa, 28.vii.1945, 1 ♂ 1 ♀, F. G. Werner lgt. (AMNH). ZAMANI- SHIMA: Ketto, xi.1945, 1 ♀, Wh. Nutting lgt. (AMNH). – CHINA: GUIZHOU: Kouy-Tchéou [= Guizhou], Rég. de Pin-Fa, 1909, 1 ♂, Pére Cavalerie lgt., P. Kment det. (MNHN). – HAINAN: Bawangling Mts., Baote env., 19°05.3′N 109°07.4′E, 415–800 m a.s.l., 7.–8.v.2011, 1 ♀, M. Fikáček, V. Kubeček & L. Li lgt. (NMPC). – TAIWAN: MIAOLI COUNTY: hills SW Sanyi, sand-steppe, 24°22′N 120°44′E, 200–500 m a.s.l., 27.x.2008, 1 ♂, L. Dembický lgt. (MMBC). – NANTOU COUNTY: Meifeng, 24°05′25″N 121°10′21″E, 10.–11.v.2001, 1 ♂ 1 ♀, K. Schönitzer lgt. (ZSMC); Puli-Yüchin, Sun Moon Lake, 16.–18.vi.1993, 2 ♂♂, J. Dalihod lgt. (ZJPC); road No. 14, NE Puli Reyen, Shi-Reg. Meifeng, Tamblingan Lakes, Nat. Res., 24°06′N 121°10′E, 2.200 m a.s.l., 9.–11.xi.2002, 1 ♀, W. Schlacht lgt. (ZSMC). – NEW TAIPEI CITY: Shiuesham Mts., Fusham Botanical Garden, ca. 24°46′N 121°35′E, 650 m a.s.l., 17.–22.vi.2000, 1 ♂ 1 ♀, W. Schlacht lgt. (ZSMC). – TAITUNG COUNTY: 14 km W of Chihshang, 29°09′N 121°04′E, 900 m a.s.l., 16.xi.2008, 30 ♂♂ 32 ♀♀, L. Dembický lgt. (MMBC, NMPC). – AFGHANISTAN: KABUL: Sarobi [= Sarowbi or Surobi], 1100 m a.s.l., 16.x.1961, 2 ♀♀, G. Ebert lgt., L. Hoberlandt 1984 det. (NMPC). NURISTAN: Bashgultal [= Landaisin river aka Bashgul river], 1300 m a.s.l., 24.v.1953, 1♀, J. Klapperich lgt., L. Hoberlandt 1984 det. (NMPC). – PAKISTAN: PUNJAB: Murree Hills, Bhurban, 1,900 m a.s.l., 3.–6.v.1978, 1♂, C. Holzschuh lgt. (EHIA); Murree, no date, 2 ♀♀, coll. Atkinson (BMNH). – INDIA: HIMACHAL PRADESH: Simla [= Shimla], 3.–4. ix. 1975, 2150 m a.s.l., 2 ♀♀, A. Čejchan & L. Hoberlandt lgt. (NMPC). – WEST BENGAL: Boutan, Pedong, 1897, 2 ♀♀, coll. Oberthür, P. Kment det. (MNHN). – NEPAL: BAGMATI ZONE: Kathmandu, British Embassy, 4500´[= 1372 m a.s.l.], 20.v.– 23.vi.1983, 2 ♀♀, M. J. D. Brendell lgt. (BMNH); Nagarjun, 27°45′N 85°17′E, l, 387 m a.s.l., forest, 29.vii.2000, 1 ♀, J. Schneider lgt. (MMBC). KARNALI ZONE: Rara Lake, 3300 m a.s.l., 6.–10.ix.1984, 2 ♀♀, Jt. SVC Rara Exped., M. G. Allen lgt. (BMNH). – KOSI ZONE: Kuwapani-Chichlia, 2,100–2,300 m a.s.l., 24.iv.1984, 1 ♂, I. Löbl & A. Smetana lgt. ( MHNG). – BHUTAN: Bhoutan Anglais, 1900, 3 ♂♂ 2 ♀♀, coll. Oberthür, P. Kment det. (MNHN); Maria Basti, 1897, 1 ♂ 1 ♀, coll. Oberthür, P. Kment det. (MNHN). – SRI LANKA: Kandy env., 600 m a.s.l., 1.– 18.iv.1991, 2 ♂♂ 1 ♀, J. Kolibáč lgt. (ZSMC). – MYANMAR: KAYIN: Dawna [Range], 10.v.1992, 1 ♂, S. Steinke lgt. (ZMHB). – TENASERIM: Tenass Vall, Myitta, no date, 1 ♀, Doherty lgt., coll. Distant, P. Kment det. (BMNH). – YANGON: Rangoun [= Yangon], 1902, 1 ♂, J. Claine lgt., P. Kment det. (MNHN). – VIETNAM: AN GIANG PROVINCE: Chaadoc [= Chau Doc], 1898, 1 ♂, coll. Noualhier (MNHN). – BAC GIANG PROVINCE: Dong Lo, no date, 1 ♂, no collector, P. Kment det. (MNHN). – BAC THAI PROVINCE: Quang Chu, 25.v.1987, 1 ♀, Mats.-Dl.-Top. lgt. (HNHM). – CAO BANG PROVINCIE: Phia Den, 8.viii.2010, light trap, 1 ♀, J. Constant & P. Limbourg lgt. (ISNB); Pia Oac Mt. Nat. Res., 22°36′N 105°53′E, pine forest, 9.viii.2010, light trap, 18♂♂ 8♀♀, J. Constant & P. Limbourg lgt. (ISNB); ditto, 4.–6.viii.2010, light trap, 8 ♂♂ 9 ♀♀, J. Constant & P. Limbourg lgt. (ISNB). – HANOI PROVINCE: Hanoi, ii.1986, 1 ♂ 1 ♀, P. Valíček lgt. (ZJPC); Hanoi, Trau Quy – Gia Lam, I.G. 31.508, 28.xi.2004, 1 ♀, X. Lammens lgt. (ISNB). – HOA BINH PROVINCE: Tonkin, Hoabinh, i.1917, 1 ♂, R. V. de Salvaza lgt. (BMNH). – LAO CAI PROVINCE: Sa Pa, v.1990, 3 ♂♂ 2♀♀, J. Picka lgt. (NMPC); ditto, vi.1990, 4♂♂ 6 ♀♀, Duong Tat Tu lgt. (NMPC); 16 km west of Sa Pa, frontier base camp, 1,800 m a.s.l., 16.iii.1989, 3♂♂, L.Peregovits & T.Vásárhelyi lgt. (HNHM). – NGHE AN PROVINCE: forestière [= gamekeeper’s lodge] Quy Chau, forest pluv. trop. semidecid. [= tropical semideciduous rain forest], 200 m a.s.l., at light, 24.viii.1963, 1 ♀, T. Pocs lgt. (HNHM). – NINH BINH PROVINCE: Cuc Phuong, 3.–10.v.1966, 1 ♂, Gy. Topál lgt. (HNHM). – QUANG TRI PROVINCE: Da Krong Nat. Park, 16°37′N 106°47′E, I.G. 31.933, 5.–10.vii.2011, light trap, 6 ♀♀, J. Constant & J. Bresseel lgt. (ISNB). – TUA THIEN- HUE PROVINCE: Hue, 5.iv.1983, 1 ♀, Macháček lgt. (NMPC). – VINH PHU PROVINCE: Me Linh Station for Biodiversity, 30.–31.v.2007, on light, 1 ♂, P. Grotaert lgt. (ISNB); Tam Dao, 900 m a.s.l., 27.v.–2.vi.1986, 4 ♂♂ 2 ♀♀, J. Horák & V. Švihla lgt.; 17.–21.iii.1990, 1 ♂ 1 ♀, J. Horák lgt. (NMPC); ditto, 1,200 m a.s.l., l 3.x.1986, 1 ♂, T. Vásárhelyi lgt. (HNHM); ditto, 9.v.1987, 1 ♂, Natskási & Topál lgt. (HNHM); ditto, 200 m a.s.l., 12.x.1988, 1 ♂, T. Vásárhelyi lgt. (HNHM); ditto, xi.1990, 2 ♀♀, G. Csorba lgt. (HNHM); ditto, I.G. 31.668, 25.–28.viii.20l0, day catch, 1 ♀, 25.–26. viii.2010, at light, 1 ♀, J. Constant & P. Limbourg lgt. (ISNB); ditto, 25.–28.viii.2010, light trap, 1 ♀, J. Constant & P. Limbourg lgt. (ISNB); ditto, 21°31′N 105°33′E, I.G. 31.933, 25.–30.vii.2011, 1 ♀, J. Constant & J. Bresseel lgt. (ISNB); ditto, 21°31′N 105°33′E, 30.vii.2011, at light, 1 ♀, J. Constant & J. Bresseel lgt. (ISNB); Tonkin, Guang Yen [= Quang Yen], 26.iii.1915, 1 ♀, R.V. de Salvaza lgt. (BMNH). – LAOS: ATTAPEU PROVINCE: Annam Highlands Mts., Dong Ampan NBCA, Nong Fa (crater lake) env., 15°05.9′N 107°25.6′E, ca. 1,160 m a.s.l., 30.iv.–4.v.2010, 1 ♂ 1 ♀, J. Hájek lgt. (NMPC). – CHAMPASAK PROVINCE: Bolavens Plateau, ca. 3 km SE Ban Lak 40 (village), coffea plantation / forest, 15°09.8′N 106°08.5′E, 1070 m a.s.l., 9. v.2010, 2 ♂♂, J. Hájek lgt. (NMPC). – HUA PHAN PROVINCE: Ban Saluei → Phou Pane Mt., 20°12–13.5′N 103°59.5′–104°01′E, 1340–1870 m a.s.l., 15.iv.–15.v.2008, Lao collectors lgt. (NMPC); Phu Phan Mt., 20°12′N 104°01′E, 1,500–1,900 m a.s.l., 17.v.–3.vi.2007, 4 ♂♂ 5 ♀♀, V. Kubáň lgt. (MMBC). – SEKONG PROVINCE: ca. 12 km S Sekong, Tad Faek waterfalls, 15°14.7′N 106°45.l′ E, 118 m a.s.l., at light, 8.–12.v.2010, 1 ♂, J. Hájek lgt. (NMPC). – VIENTIANE PROVINCE: Lao Pako env., 55 km NE of Vientiane, 200 m a.s.l., 1.–4.v.2005, 3 ♂♂ 3 ♀♀, J. Bezděk lgt., Z. Jindra det. (NMPC); Kamphaeng Nakhon Viang Chan prov., 25 km of Vientiane, 1 ♀, Boudon lgt. (MNHN). – CAMBODIA: KAMPOT PROVINCE: Bokor Nat. Park, Hill Station, scrub, 22.iv.2005, 1 ♀, K. Smets & I. Var lgt. (ISNB). – PURSAT PROVINCE: Phnom Samkos W.S., Temple Hill, grassland, bamboo forest, 18.v.2005, light trap, 1♂, K. Smets & I. Var lgt. (ISNB). – SIEM REAP PROVINCE: Angkor Thorn, 5.iii.2005, light trap, 1 ♂, D. R. Jump lgt. (ISNB); ditto, viii.2005, by net, 1 ♂, D. R. Jump lgt. (ISNB). – THAILAND: KAMPAENG PHET PROVINCE: Khiong Lan Nat. Park, 16°07′N 99°16′E, UV pan trap, 6.iv.2003, 2 ♀♀, Sites, Vitheepradit, Prommi & Setaphan lgt. (MUUS). – LOEI PROVINCE: Na Haeo, Biol. Station, light trap, 5.–l 2.v.2001, 4 ♂♂, J. Constant & P. Grootaert lgt. (ISNB); ditto, Na Haeo, river bank, on light trap, l 5.v.2003, 1♂, P. Grootaert, J. Constant & K. Smets lgt. (ISNB); Na Haeo, forest clearing, on light trap, 16.v.2003, 1 ♂, J. Constant & K. Smets lgt. (ISNB). – MAE HONG SON PROVINCE: Northern – Pai, 1.–12.v.2001, 1 ♂ 1 ♀, R. Kocina lgt., Z. Jindra det. (NMPC); Namtok Mae Surin Nat. Park, Ma Nam Pai, 19°21′N 97°59′E, 310 m a.s.l., 31.iii.2003, 1 ♂ 1 ♀, UMC & CMU teams lgt. (MUUS); Pang Mapha, 190°31.159′N 98°14.844′E, 19.iii.2009, 1 ♀, R.V. Sites lgt. (MUUS). – NAN PROVINCE: Mae Charin Nat. Park, 18°36′N 100°59′E, 285 m a.s.l., UV pan trap, 22.iv.2003, 1 ♂ 1 ♀, Vitheepradit, Prommi & Setaphan lgt. (MUUS). – PHRAE PROVINCE: Wieng Ko Sai Nat. Park, 17°58′N 99°35′E, 350 m a.s.l., pan UV light trap, 29.iii.2003, 1 ♀, Sites, Vitheepredit & Prommi lgt. (MUUS). – PITHSANULOG PROVINCE: Phu Hin Rongkhla Nat. Park, on vapor light at guest house, 11.iii.2002, 3 ♂♂ 4 ♀♀, Sites, Vitheepredit & Kirawanich lgt. (MUUS); ditto, Phu Hin Rongkhla Nat. Park, Nam Tok Romglao, 16°59′N 101°00′E, 1,190 m a.s.l., 22.iv.2002, 1 ♂, CMU Team lgt. (MUUS); ditto, Phu Hin Rongkhla Nat. Park, 17°00′N 100°59′E, 1,144 m a.s.l., trap at guest house, 5.v.2003, 1 ♀, Vitheepradit, Prommi & Ferro lgt. (MUUS); ditto, Thung Salaeng Luang, 12.v.2004, 1 ♀, Vitheepradit & Prommi lgt. (MUUS). – PROCHUAP KHIRI KHAN PROVINCE: Amphur Kui Buri, Forest, Plantation Station, on black light, 12°04′N 99°43′E, 147 m a.s.l., 22.iv.2002, 1 ♀, Vitheepradit & Kirawanich lgt. (MUUS). – SURATTHANI PROVINCE: Suratthani env., 1989, 1 ♂, S. Lehman lgt. (ZSMC). – UPPER RATHATARU PROVINCE: Phu Chong Na Yoi Nat. Park, 14°26.l25′N 105°l5.360′E., 1 ♀, Sites & Vitheepradit lgt. (MUUS); ditto, Phu Chong, Nat. Park, Kaeng Ka Loa, 14°26.178′N 105°16.831′E, 182 m a.s.l., 11.iv.2004, 1 ♀, Sites & Vitheepradit lgt. (MUUS). – MALAYSIA: MALAY PENINSULA: FEDERAL TERRITOTY OF KUALA LUMPUR: Kuala Lumpur, at light, 3.xi.1922, 1 spec., ex F.M.S. Museum, P. Kment det. (BMNH). – KEDAH: : West Coast, Langkawi Island, 28.iv.1928, 1 ♂, 29.iv.1928, 1 ♀, ex F.M.S. Museum, P. Kment det. (BMNH). – PAHANG: Cameron’s Highlands, 4,000 –5,000 ft [= 1219–1524 m a.s.l.], 10.vi.1935, H. M. Pendlebury lgt., P. Kment det. (BMNH). – PERAK: Taiping, x.1977, 3 ♂♂ 7 ♀♀, P. Pfanner lgt. (MHNG). – SELANGOR: Bukit Kutu, 3,500 ft [= 1067 m a.s.l.], 6.ix.1929, 1 ♀, 13.ix.1929, 1 ♀, 14.ix.1929, 1 ♀, H. M. Pendlebury lgt., P. Kment det. (BMNH). – KALIMANTAN: SABAH: Kinabalu, xi.1978, 1 ♂, no collector (MNHN) ; Mt. Kinabalu, Nat. Park, 1,500 m a.s.l., 22.–27.iv.1996, 1 ♀, S. Snäll lgt. (ZJPC). – SARAWAK: Gunong Mulu Nat. Park, RGS. Mulu Exped., 1 ♀, J. D. Holloway lgt. (BMNH). – STATE NOT IDENTIFIED: North Borneo, Tawan, on cocoa, 13.ii.1959, 3 ♂♂, K. L. Leong lgt., M. S. K. Ghauri 1963 det. (BMNH). – SINGAPORE: Singapore, 89-38, no date, 1 spec., no collecetor (BMNH); Singapore, 96-10, no collector, 1 ♂ 1 ♀, P. Kment det. (BMNH); Singapore, 97-12, 1 ♀, H. N. Ridley lgt., P. Kment det. (BMNH); Singapore, 1900-114, H. N. Ridley lgt., P. Kment det. (BMNH). – BRUNEI: Temburong District, ridge NE Kuala Belalong, 125 W m. v. light , ca. 300 m a.s.l., xi.1992, 1 ♂ 2 ♀♀, J. H. Martin lgt., P. Kment det. (BMNH); S[ungai] Selanjak, 4463.1432, mangrove, 0 m a.s.l., 6.–7.iii.1984, 1 ♂, Helps T. P. G. Maj. [lgt.], P. Kment det. (BMNH). – INDONESIA: BALI: Beduqul distr., 1,300 m a.s.l., i.–ii.2005, 1 ♂♂ 2 ♀♀, S. Jákl lgt. (ZJPC). – JAVA: JAWA BARAT PROVINCE: Bibidlijan, 10.xi.1937, 3 ♂♂ 2 ♀♀, M. E. Walsch lgt. (ZMHB) ; Bogor, no date, 1 ♂, Handschin lgt. (ZMHB); Radjamandala, 1,200 m a.s.l., 6.xi.1937, 4 ♂♂, M. E. Walsch lgt. (ZMHB); Soekaboemi [= Sukabumi], 1 ♀, F. Weber lgt. (ZMHB). – JAWA TIMUR PROVINCE: Meru, Betiri Nat. Park, Malangsari villge, 20.v.–5.vi.1996, 1 ♂, S. Jákl lgt. (ZJPC). – KALIMANTAN: KILIMANTAN SELATAN PROVINCE: S. Kalimantan prov., Loksado 17 km NE, I.G. 31.175, 2 ♀♀, St. Jákl lgt. (ISNB). – KILIMANTAN TIMUR PROVINCE: Balikpapan, 1947/1948, 1 ♀, W. A. Mohler lgt. (ZMHB). – MENTAWAI ISLANDS: Nias Island, East coast, Lawlo, 26.ix.1979, 1 ♂, Ebner lgt. (EHIA). – SUMATRA: RIAU PROVINCE: Indragiri, 1899, 1 ♀, A. Mechel lgt. (ZMHB) . – SUMATERA BARAT PROVINCE: Bukittinggi, hotel ‘Bukit Tinggi View’, 00°15′30″N 100°21′13″E, 980 m a.s.l., 19.–23.ii.2002, 1 ♀, T. Kothe lgt. (ZSMC); Mt. Singgalang, Annai valley env., 600 m a.s.l., v.2006, 12 ♂♂ 14 ♀♀, S. Jákl lgt. (ISNB, ZJPC). – SUMATERA UTARA PROVINCE: Dolak Ulu, Kora Kora, 20.ix.1979, 1 ♂ 1 ♀, Ebner lgt. (EHIA); Sidikalang, 02°41′51″N 98°18′18″E, 1,250 m a.s.l., 15.–17.ii.2002, 1 ♂ 1 ♀, T. Kothe lgt. (ZSMC); near Sipirok, Danau Marsaput, 01°37′50″N 99°20′30″E, 1,435 m a.s.l., 1 ♀, T. Kothe lgt. (ZSMC) ; S. Pomatang Siantar, 1,100 m a.s.l., 28.–29.x.1992, 1 ♀, E. de Bros lgt. (ZMHB). – PHILIPPINES: LUZON: IFUGAO PROVINCE: Mayoyao, viii.1988, 1 ♂ 1 ♀, P. Lays lgt. (ISNB). ILICOS NORTE PROVINCE: E of Salsona, 18°05′N 120°54′E, 1,450 m a.s.l., 31.iii.2000, 2 ♀♀, L. Dembický lgt. (MMBC) . – MINDANAO: SOUTH COTOBATO PROVINCE: Lake Sebu, Mt.Tasaday, 6°18′N 124°42′E, v.–vi.1994, 1 ♂, P. Lays lgt. (ISNB) ; Lake Sebu, 6°13′N 124°42′E, 700 m a.s.l., at light trap (neon), 1.–15.i.1998, 1 ♂ 2 ♀♀, P. Lays lgt. (ISNB) . LANAO DEL NORTE PROVINCE: Iligan, 1 ♀, W. Schulze lgt. (ZSMC) . – BUKIDNON PROVINCE: Bukidnon, 1.–10.x.1988, 3 ♂♂ 1 ♀, K. Černý lgt. (MMBC). – NEGROS ISLAND: NEGROS ORIENTAL PROVINCE: Camp Lookout, Dumaguete, 15.ii.–15.iv.1961, 1 ♂, T. Schneiria & A. Reyes lgt. (AMNH). – PALAWAN: Irawan Forest, 1000 ft, swept, B.M. 1988-2, i.–ii. 1988, 1 ♀, J. H. Marting lgt. (BMNH); Mantalingajan Range, Pinigisan, approx. 600 m a.s.l., 31.viii.1961, 1 ♂, 8.ix.1961, 1 ♂, 9.ix.1961, 2 ♂♂ 2 ♀♀, all Noona Dan Expedition (ZMUC); Pa1awan, Irawan Forest, swept, 1,000 ft [= 305 m a.s.l.], i.–ii.1988, 1 ♀, J. H. Marhn lgt. (BMNH). – TAWI TAWI ISLANDS: Tarawakan, north of Batu Batu, 14.xi.1961, 3 ♀♀, Noona Dan Expedition (ZMUC); ditto, Lapid Lapid at Manalik Channel, 19.xi.1961, 1 ♀, Noona Dan Expedition (ZMUC).

Distribution. Japan (Honshu, Kyushu, Ryukyus) ( SCOTT 1874, DISTANT 1883, UHLER 1897, BLÖTE 1931, MANNA et al. 1985, KERZHNER 2001, KOHNO et al. 2002, TOMOKUNI 2006; this paper), Korea (South, Chejudo Island) ( LEE & KWON 1991, KWON et al. 2001, KERZHNER 2001), China (Fujian, Guangdong, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan, Xizang, Zhejiang, Hainan) ( STÅL 1870, WALKER 1873, DISTANT 1903a, LIU 1981, HUA 2000, KERZHNER 2001; this paper), Taiwan ( ESAKI 1926, BLÖTE 1931, KERZHNER 2001, RÉDEI et al. 2009; this paper), Afghanistan ( KERZHNER 2001, no details; this paper), Pakistan (Khyber Pakhtunkhwa, Punjab) ( AHMAD & ABBAS 1987; this paper), India (Andhra Pradesh, Arunachal Pradesh, Assam, Bihar, Haryana, Himachal Pradesh, Jammu and Kashmir, Karnataka, Madhya Pradesh, Meghalaya, Tamil Nadu, Uttarakhand, West Bengal) ( DISTANT 1879a, 1903a; MAXWELL- LEFROY 1909; SCHMIDT 1931; MANNA et al. 1985; ZAMAL & CHOPRA 1990; CHAKRABARTY et al. 1994; SAHA & BAL 2007; STEHLÍK 2007a; CHANDRA et al. 2012; CHANDRA & KUSHWAHA 2013; SINGH & BANYAL 2013; this paper), Bhutan (new record), Nepal ( STEHLÍK 2004; this paper), Sri Lanka ( WALKER 1873, KIRBY 1891; this paper), Bangladesh ( WALKER 1873), Myanmar ( DISTANT 1879b, 1903a; this paper), Vietnam (new record), Laos ( STEHLÍK 2005a; this paper), Cambodia (new record), Thailand ( DISTANT 1903b, STEHLÍK & JINDRA 2003; this paper), Malaysia (Peninsular Malaysia: Federal Territory of Kuala Lumpur, Kedah, Pahang, Penang, Selangor; Kalimantan: Sabah, Sarawak) ( WALKER 1873, BREDDIN 1900; DISTANT 1903b; this paper), Singapore (new record), Brunei (new record), Indonesia (Bali, Java, Kalimantan, Mentawai Islands: Nias, Sumatra) ( BURMEISTER 1834, 1835; STÅL 1855, as Ph. bimaculata, 1870 ; WALKER 1873; LETHIERRY 1888; BREDDIN 1901b; DISTANT 1903a; BLÖTE 1931; SCHMIDT 1931), Philippines (Basilan, Luzon, Mindanao, Negros, Palawan, Tawi Tawi Island) ( BURMEISTER 1834, STÅL 1870, WALKER 1873, TAEUBER 1927; this paper).

Distributional records from Australia and Australian Region by BREDDIN (1900, 1901a), CASSIS & GROSS (2002), and VOIGT (2006) belong to Ph. gutta famelica (see e.g., STEHLÍK 2007a). The records of Ph. gutta from Sulawesi ( BREDDIN 1901a), Timor ( Blöte 1931), and Halmahera ( BREDDIN 1900) apparently belong to Ph. gutta famelica as well. The record of Ph. gutta famelica from China: Sichuan by VOIGT (2006) represents a misidentification of Ph. gutta gutta .

Figures

Figs 17–19. Physopelta (Neophysopelta) gutta gutta (Burmeister, 1834), male. 17 – fore coxae and trochanters (magnification 50×), 18 – detail of procoxa and protrochanter (95×), 19 – ventral surface of profemur (37×). Scale bars: 0.5 mm. (SEM micrographs: P. Kment).

Figs 20–25. Physopelta (Neophysopelta) gutta gutta (Burmeister, 1834), female. 20 – meso- and metapleuron with the external scent efferent system (magnification 30×); 21 – detail of ostiole (300×); 22 – ostiole, peritreme, and surrounding evaporatorium (130×); 23 – apical portion of peritreme in detail (300×); 24 – evaporatorium, detail of mycoid microsculpture mesad of the median furrow (1200×); 25 – peritreme, detail of microsculpture (smooth peritremal surface of median furrow left, spinules of the remaining peritremal surface right; 1200×). Lettering: ev – evaporatorium, mf – median furrow of peritreme, o – ostiole, pe – peritreme. Scale bars: 10 μm (Figs 24, 25), 50 μm (Figs 21, 23), 100 μm (Fig. 22), 500 μm (Fig. 20). (SEM micrographs: P. Kment).

Figs 26–29. Physopelta (Neophysopelta) gutta gutta (Burmeister, 1834), male, pygophore (magnification 50×): 26 – dorsal view, 27 – postero-dorsal view, 28 – posterior view, 29 – lateral view. Scale bars: 0.5 mm. (SEM micrographs: P. Kment).

Figs 30–34. Physopelta (Neophysopelta) gutta gutta (Burmeister, 1834), male, paramere (different orientations, magnification 190×). Scale bars: 0.2 mm. (SEM micrographs: P. Kment).

Figs 93–98. Kmentia festiva (Fabricius, 1803). 93 – female; 94 – male; 95–96 – fore leg (95 – large male, 96 – small male); 97 – ventral surface of profemur. 98 – Physopelta (Neophysopelta) gutta gutta (Burmeister, 1834), ventral surface of profemur. (Photos: L. Dembický).