Calapnita anai , Bernhard A. Huber, 2017

Bernhard A. Huber, 2017, Revision and cladistic analysis of the Southeast Asian leaf-dwelling spider genus Calapnita Simon (Araneae, Pholcidae), Zootaxa 4219 (1), pp. 1-63: 29-30

publication ID 10.5281/zenodo.273086

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scientific name

Calapnita anai

sp. nov.

Calapnita anai  sp. nov. Figs 11–12View FIGURES 3 – 12, 107–119View FIGURES 107 – 111View FIGURES 112 – 119

Calapnita phasmoides  ” (misidentification): Huber 2011: 59 (only specimens assigned tentatively from Sumatra and Java).

Diagnosis. Males are easily distinguished from most congeners (except C. phasmoides  and C. deelemanae  ) by simple appendix with small subdistal side-branch ( Fig. 110View FIGURES 107 – 111); from C. phasmoides  and C. deelemanae  by tip of procursus (not strongly bent; indistinct retrolateral hump; dorsally gradually widening; Figs 107–109View FIGURES 107 – 111); females of these three species may be indistinguishable externally; internally, the round pore plates close together and the straight anterior margin ( Fig. 111View FIGURES 107 – 111) may be distinctive.

Etymology. The species name is derived from the type locality; noun in apposition.

Material examined. Holotype. INDONESIA-SUMATRA: ♂, ZFMK (Ar 16016), West Sumatra, Lembah Anai  Forest Reserve , forest along stream (0.472°S, 100.363°E), 600 m a.s.l., 19.x.2009 (S. Sutono).GoogleMaps 

Other material. INDONESIA-SUMATRA: 3♀, ZFMK (Ar 16016), together with holotype; 3♂ 5♀ in absolute ethanol, ZFMK (Ind 134), same data.

SINGAPORE: 2♂ 1♀, ZFMK ( Ar 16017), MacRitchie Reservoir Park (1°21.3’N, 103°48.8’E), 50 m a.s.l., on leaves, 14.ii.2015 (B.A. Huber, J.K.H. Koh, D. Court)GoogleMaps  ; 5♂ 4♀ 2 juvs in absolute ethanol, ZFMK (Mal 207), same dataGoogleMaps  . 1♀ in absolute ethanol, ZFMK ( Mal 221), Rifle Range (1°21.3’N, 103°47.8’E), 50 m a.s.l., on leaf, 15.ii.2015 (B.A. Huber, D. Court).GoogleMaps 

MALAYSIA: 2♂ 1 juv., ZFMK (Ar 16018), Pahang, Cameron Highlands (4°27.6’N, 101°23.5’E), 1200 m a.s.l., forest along ‘trail 9’, on leaves, 25.ii.2015 (B.A. Huber); 4 juvs in absolute ethanol, ZFMK (Mal 281), same dataGoogleMaps  . 1♀ in absolute ethanol, ZFMK (Mal 260), Pahang, Ulu Dong (3°56.2’N, 102°01.9’E), 190 m a.s.l., forest near river, on leaf, 21.ii.2015 (B.A. Huber, A.R.M. Ghazali, K.A. Braima).GoogleMaps 

THAILAND: 1♂ in absolute ethanol, ZFMK (Mal 307), Narathiwat, Hala Bala Wildlife Sanctuary , ‘site 3’ (5°48.5’N, 101°49.4’E), 220 m a.s.l., forest near road, leaf litter, 2.iii.2015 (B.A. Huber, B. Petcharad)GoogleMaps  . 1♀, ZFMK (Ar 16019), Narathiwat, Hala Bala Wildlife Sanctuary , ‘site 2’ (5°48.4’N, 101°48.6’E), 330 m a.s.l., forest near river, on leaves, 2.iii.2015 (B.A. Huber, B. Petcharad)GoogleMaps  . 1♀ in absolute ethanol, ZFMK (Mal 301), Hala Bala Wildlife Sanctuary , ‘site 1’, forest at river near headquarters (5°47.8’N, 101°49.9’E), 90 m a.s.l., on leaf, 1.iii.2015 (B.A. Huber, B. Petcharad)GoogleMaps  . 3♂ 2♀, ZFMK (Ar 16020), Satun, Thaleban National Park (6°42.6’N, 100°10.2’E), forest near headquarters, 110 m a.s.l., on leaves, 5.iii.2015 (B.A. Huber, B. Petcharad)GoogleMaps  ; 2♀ in absolute ethanol, ZFMK (Mal 327), same data.GoogleMaps 

Description. Male (holotype).

MEASUREMENTS. Total body length 5.6, carapace width 1.0. Leg 1: 35.9 (8.4 + 0.4 + 8.5 + 16.6 + 2.0), tibia 2: 6.1, tibia 3: 3.6, tibia 4: 5.5; tibia 1 L/d: 100. Distance PME-PME 300 µm, diameter PME 90 µm, distance PME- ALE ~30 µm; no trace of AME.

COLOR. Entire animal mostly whitish to pale ochre-yellow, patellae and tibia-metatarsus joints slightly darker brown, abdomen monochromous whitish.

BODY. Habitus as in Fig. 11View FIGURES 3 – 12; ocular area barely elevated, each triad on very low hump; carapace without median furrow; clypeus unmodified; sternum as wide as long (0.56), unmodified.

CHELICERAE. As in close relatives (cf. figs 172, 204 in Huber 2011), with pair of scaly apophyses near lamellae ( Fig. 112View FIGURES 112 – 119) and pair of distinct but whitish proximal humps; without modified hairs; without stridulatory ridges.

PALPS. In general similar to C. phasmoides  (cf. figs 194–195 in Huber 2011); tip of trochanter apophysis more strongly curved; procursus rather simple distally, gradually widening in lateral view, with indistinct retrolateral process ( Figs 107–109View FIGURES 107 – 111); appendix with small retrolateral cone between tip and subdistal side branch ( Figs 110View FIGURES 107 – 111, 115View FIGURES 112 – 119).

LEGS. Without spines and curved hairs; few vertical hairs; retrolateral trichobothrium on tibia 1 at 2.5%; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 apparently with ~30 pseudosegments but only distally fairly visible in dissecting microscope.

Male (variation). Tibia 1 in 8 other males (including “ C. phasmoides  ” male from Sumatra in Huber 2011): 7.4– 8.6 (mean 7.8).

Female. In general similar to male; eye triads slightly closer together (distance PME-PME 250 µm). Tibia 1 in 12 females (including “ C. phasmoides  ” females from Sumatra in Huber 2011): 6.1–7.2 (mean 6.7). Epigynum weakly sclerotized, internal dark structure visible through cuticle near posterior margin; with simple posterior ‘knob’ ( Fig. 119View FIGURES 112 – 119); internal genitalia as in Fig 111View FIGURES 107 – 111.

Natural history. At Cameron Highlands, all specimens were collected from banana leaves near the forest edge; no specimen was seen in the forest. Egg-sacs (seen at several localities) consisted of a single row of 9– 12 eggs ( Fig. 12View FIGURES 3 – 12). Webs consisted of a rather dense sheet of silk, but could only be seen against the light when the leaf was viewed from the side.

Distribution. Widely distributed on the Malay Peninsula and West Sumatra ( Fig. 281View FIGURE 281); the single female from Java that was tentatively assigned to C. phasmoides in Huber 2011  may in fact belong to this species.


Zoologisches Forschungsmuseum Alexander Koenig














Calapnita anai

Bernhard A. Huber 2017

Calapnita phasmoides

Huber 2011: 59