Galactodenia Sundue & Labiak, Syst. Bot.

Smith, Alan R., Kessler, Michael, León, Blanca, Almeida, Thaís Elias, Jiménez-Pérez, Iván & Lehnert, Marcus, 2018, Prodromus of a fern flora for Bolivia. XL. Polypodiaceae, Phytotaxa 354 (1), pp. 448-450 : 448-450

Galactodenia Sundue & Labiak, Syst. Bot. 37(2): 340. 2012.

Galactodenia comprises five species, all neotropical, and all but one exclusively in South America; only one species occurs in Bolivia. Two species, including G. subscabra in Bolivia, were included in Terpsichore Group 3 of Smith (1993); the remaining three species are recently described and relatively rare, narrow endemics ( Sundue et al. 2012). The genus ranges from Costa Rica to Colombia and Venezuela, south in the Andes to Bolivia. Characters of the genus include pendent fronds with thin 1-pinnate or slightly more divided blades, weakly developed hydathodes, absence of setae in all but one species, and 1-forked and 3-4-celled, translucent, tubular, glandular hairs all over the blades on both surfaces ( Sundue et al. 2010). In phylogenetic analyses, Galactodenia is most closely related, and sister, to the clade that includes Lellingeria , Melpomene , and Stenogrammitis ( Ranker et al. 2004, Lehnert et al. 2009, Labiak et al. 2010 a, Sundue et al. 2010, Sundue et al. 2012).