Ceradenia tryonorum

Smith, Alan R., Kessler, Michael, León, Blanca, Almeida, Thaís Elias, Jiménez-Pérez, Iván & Lehnert, Marcus, 2018, Prodromus of a fern flora for Bolivia. XL. Polypodiaceae, Phytotaxa 354 (1), pp. 448-450 : 448-450

Ceradenia tryonorum B. León & A.R.Sm., Amer. Fern J. 93(2): 81(–84), f. 1. 2003.

Range: — Peru and Bolivia (LP).

Ecology: —Rare, known only from a single Bolivian specimen: Jiménez 5571 (LPB); epiphyte; 3050 m.

Notes: — León and Smith (2003) described this species from a single Peruvian collection, and the species has subsequently been discovered in Bolivia. This particular pattern exemplifies the spotty, seemingly rare nature of distributions of many Andean mostly epiphytic grammitids, nearly all found in primary, undisturbed montane forests; however, it is probably also true that this group is overlooked by general collectors because of their inconspicuous, sometimes confusingly similar morphology.

Ceradenia tryonorum is very unusual within the genus in having hydathodes present on the adaxial blade surface. Generally the anhydathodous condition is the rule in Ceradenia and the closely related genera Enterosora and Zygophlebia , but hydathodes are found also in Enterosora asplenioides L.E.Bishop , from Colombia and Ecuador. This species also has linear sori, and these two shared characters in both the Enterosora and Ceradenia tryonorum may reflect deeper relationships between Ceradenia and Enterosora than their current generic placement suggests ( León & Smith 2003).