Isaraselis cladiva, Drohojowska, Jowita & Szwedo, Jacek, 2013

Drohojowska, Jowita & Szwedo, Jacek, 2013, The first Aleyrodidae from the Lowermost Eocene Oise amber (Hemiptera: Sternorrhyncha), Zootaxa 3636 (2), pp. 319-347 : 336-337

Isaraselis cladiva sp. n.

( Figures 58 –72)

Diagnosis. As for genus: claspers elongate, longer than pygofer, distinctly tapering toward apex, crossed at apex, with distinct basal teeth.

Description. Measurements given in Table 1 . Head massive, placed in axis with the body. Compound eyes distinctly divided into upper and lower portions, width of lower portion less than half that of upper portion. Antennae 8-segmented. Scape as long as wide, pedicel huge, distinctly widened in apical portion, about 2.5× as long as wide, antennal segment III twice as long as pedicel, segments VI and VII equal in length, VIII tapering apically beginning at half of its length. Apical segment of rostrum tapering, slightly shorter than preapical one. Rostrum reaching hind leg coxae. Fore wings 2.5× as long as wide, widest at ½ of its length, costal margin merely curved at base, then straight, anterior angle very widely rounded, apical margin rounded, posteroapical angle rounded, posterior margin straight, all margins tuberculate, tubercles asetose. Apex of clavus at half of fore wing length; postclaval margin straight. Common stem Sc+R+CuA thick, short, stems Sc+R and CuA diverging at acute angle; stem Sc+R straight, forked at half of fore wing length; branch R1 not reaching costal margin, branch Rs almost straight reaching apical margin; stem CuA straight, slightly curved at apical portion, not reaching posteroapical angle; claval vein CuP distinct. Hind wing very narrow at base, widest at half of its length; costal margin straight, anteroapical angle very widely rounded, apical margin rounded, posteroapical angle widely rounded, posterior margin straight. Costal margin up to half of hind wing length covered with tubercles bearing short, evenly dispersed spinules, the remaining margins covered with asetose tubercles. Stem Sc+R straight, branch R1 forked basad of half of hind wing length, straight, not reaching costal margin; branch Rs straight, as prolongation of stem Sc+R, not reaching apical margin; stem CuA merely curved, not reaching posteroapical angle. Hind tibia with a row of delicate setae forming a comb, in basal portion; apical portion with a few stronger setae; tarsal claw distinct, paronychium not visible. Abdomen with three distinct wax-plates, on the abdominal segments II, III and IV. Wax plates of segments II and III elongate, wider in dorsal portion, wax-plate of segment III rd the largest, wax-plate of segment IV about half of size of wax-plates of preceding segments, ellipsoidal. Pygofer tubulate, with posterior margin shallowly concave. Claspers longer than pygofer, with basal teeth, tapering apicad and curved at tips, crossed at apex.

Etymology. Specific epithet is derived from the Celtic word ‘cladivos’ meaning glaive and refers to the shape of claspers.

Age and occurrence. Lowermost Eocene, c. 55−53 Ma, Ypresian (Sparnacian), level MP7 of the mammal fauna of Dormaal (Feugueur 1963; De Ploëg et al. 1998; Nel et al. 1999, Cavagnetto 2000, De Franceschi & De Ploëg 2003, Nel et al. 2004, Nel & Brasero 2010). Department Oise, France, 43°N, 2.7°E; palaeocoordinates 44.8°N, 0.1°E; palaeocoordinates based on nearby landmark (Paleobiology Database 2013).

Material. Holotype, male. MNHN PA 16445. Thorax covered with bubble of gas.

Figures

FIGURES 58 – 62. Isaraselis cladiva gen. et sp. n. 58 — body, lateral view; 59 — antenna; 60 — hind leg; 61 — fore wing; 62 — hind wing.