Oedothorax paracymbialis, TANASEVITCH, 2015 INCERTAE

Lin, Shou-Wang, Lopardo, Lara & Uhl, Gabriele, 2022, Evolution of nuptial-gift-related male prosomal structures: taxonomic revision and cladistic analysis of the genus Oedothorax (Araneae: Linyphiidae: Erigoninae), Zoological Journal of the Linnean Society 195, pp. 417-584 : 552-553

‘OEDOTHORAX’ PARACYMBIALIS TANASEVITCH, 2015 INCERTAE SEDIS

( FIGS 7C , 67D , 68D , 79 )

Oedothorax paracymbialis Tanasevitch, 2015: 389 , figs 70–74 (Dm).

Type material: Holotype: India: Madras , Nilgiri , Hulical near Coonoor, right bank of Coonoor River, 1600 m, forest in ravin, sifting, ♂ 22.xi.1972, leg. C. Besuchet & I. Löbl ( MHNG, examined) .

Diagnosis:

Males: This species can be diagnosed by the slightly elevated postocular region, the prominent TPS and the large distal part of paracymbium.

Description:

Male (holotype): Total length: 1.77. Prosoma: 0.77 long, 0.63 wide, postocular region slightly elevated, with curved seta directed anteriorly ( Fig. 7C ). Eyes: AME-AME: 0.031, AME width: 0.04, AME-ALE: 0.03, ALE width: 0.08, ALE-PLE: 0.01, PLE width: 0.08, PLE-PME: 0.03, PME width: 0.07, PME-PME: 0.06. Clypeus: not hirsute. Sternum: 0.48 long, 0.49 wide. Chelicerae: mastidia absent; stridulatory striae rows widely and evenly spaced ( Fig. 67D ). Legs: Tm I: 0.62. All metatarsi with trichobothrium. Pedipalp: patella prolateral proximal vertical macrosetae absent; tibia with one prolateral, two retrolateral trichobothria; TPS striped, straight, apical-retrolaterally pointed, wide at base; TPA slightly elevated; TRA absent ( Fig. 79C ); PC large, base not visible from dorsal view, distal setae close to distal clasp, setae-bearing region greatly enlarged, distal clasp without striae, clasp directed slightly apically ( Fig. 79A ); T without papillae; PT short, without papillae, distal rim thin and smooth at margin; TS absent ( Fig. 79D ); MSA present; DSA straight, tip round; EM absent; ARP horizontally flat, highly sclerotized; LER small, without striae, not extended dorsal to E; VRP absent; TP slender; E retrolaterally spiral, middle part wide and striped ( Fig. 79E ). Opisthosoma: dorsal pattern see Fig. 68D ; PMS with mAP, two AC; PLS with triad, three AC ( Fig. 79F ).

Female: Unknown.

Distribution: Only known from the type locality.

Habitat: Forest litter.

Remarks: According to the results of our phylogenetic analysis, and the lack of the distinctive characteristics of Oedothorax s.s., this species is more closely related to species in Clade 16 that, as this species, are distributed in the Oriental realm. We provisionally leave the taxonomic status of this species unchanged until further data become available.

Figures

Figure 7. Male prosomal morphology, lateral.A, Emertongone montifera (Emerton, 1882) (Paquin & Dupérré, 2003, fig. 1187). B, Shaanxinus mingchihensis Lin, 2019 (traced from photograph). C, ‘Oedothorax’ paracymbialis Tanasevitch, 2015 (traced from photograph). D, ‘Oe.’ kodaikanal Tanasevitch, 2015 (traced from photograph). E, ‘Oe.’ cunur Tanasevitch, 2015 (traced from photograph). F, ‘Oe.’ stylus Tanasevitch, 2015 (traced from photograph). G, Atypena cirrifrons (Heimer, 1984) (traced from photograph). H, A. formosana (Oi, 1977) (Oi, 1977, fig. 2). I, ‘Oe.’ nazareti Scharff, 1989 (Scharff, 1989: fig. 7). J, Gongylidium rufipes (Linnaeus, 1758) (traced from photograph). K, Ummeliata insecticeps (Bösenberg & Strand, 1906) (Tu & Li 2004: fig. 8A). L, U. esyunini (Zhang, Zhang & Yu, 2003) (Zhang et al., 2003, fig. 2A). M, Hylyphantes graminicola (Sundevall, 1830) (traced from photograph). N, Tmeticus tolli Kulczyński, 1908 (traced from photograph). O, Oedothorax gibbosus (Blackwall, 1841) [Roberts, 1987: fig. 21 (e)]. P, Oe. trilobatus (Banks, 1896) (Bishop & Crosby, 1935: fig. 79). Q, Oe. gibbifer (Kulczyński, 1882) (Heimer & Nentwig, 1991: fig.604.5).R, Oe.apicatus (Blackwall, 1850) [Roberts, 1987: fig.23 (d)].S, Oe. retusus (Westring, 1851) [Roberts, 1987: fig. 23 (c)]. T, Oe. paludigena Simon, 1926 [Millidge, 1975: fig. (5)]. U, Oe. meridionalis Tanasevitch, 1987 (Tanasevitch, 1987: fig. 107).V, Oe. agrestis (Blackwall, 1853) (Roberts, 1987: fig. 23B).W, Oe. fuscus (Blackwall, 1834) (Roberts, 1987: fig. 23A). X, Oe. tingitanus (Simon, 1884) (Denis, 1968: fig. 11).

Figure 67. Male chelicerae, stridulatory striae on lateral side. A, Jilinus hulongensis (Zhu & Wen, 1980). B, Cornitibia simplicithorax (Tanasevitch, 1998). C, Emertongone montifera (Emerton, 1882). D, ‘Oedothorax’ paracymbialis Tanasevitch, 2015. E, ‘Oe.’ kodaikanal Tanasevitch, 2015. F, ‘Oe.’ cunur Tanasevitch, 2015. G, ‘Oe.’ stylus Tanasevitch, 2015. H, Atypena cirrifrons (Heimer, 1984). I, A. formosana (Oi, 1977). J, ‘Oe.’ nazareti Scharff, 1989. K, Gongylidium rufipes (Linnaeus, 1758). L, Ummeliata insecticeps (Bösenberg & Strand, 1906). M, U. esyunini (Zhang, Zhang & Yu, 2003). N, Hylyphantes graminicola (Sundevall, 1830). O, Tmeticus tolli Kulczyński, 1908. Scale bars 0.05 mm.

Figure 68. Male opisthosoma, dorsal view. A, Jilinus hulongensis (Zhu & Wen, 1980). B, Cornitibia simplicithorax (Tanasevitch, 1998). C, Emertongone montifera (Emerton, 1882). D, ‘Oedothorax’ paracymbialis Tanasevitch, 2015. E, ‘Oe.’ kodaikanal Tanasevitch, 2015. F, ‘Oe.’ cunur Tanasevitch, 2015. G, ‘Oe.’ stylus Tanasevitch, 2015. H, Atypena cirrifrons (Heimer, 1984). I, A. formosana (Oi, 1977). J, ‘Oe.’ nazareti Scharff, 1989. K, Gongylidium rufipes (Linnaeus, 1758). L, Ummeliata insecticeps (Bösenberg & Strand, 1906). M, U. esyunini (Zhang, Zhang & Yu, 2003). N, Hylyphantes graminicola (Sundevall, 1830). O, Tmeticus tolli Kulczyński, 1908. Scale bars 0.1 mm.

Figure 79. ‘Oedothorax’ paracymbialis Tanasevitch, 2015. A–E, male right palp, images flipped horizontally.A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, embolic division, prolateral view. F, male spinnerets. Scale bars 0.1 mm.