Ambonga apicenotata Melichar

Andrew Hamilton, K. G., 2014, The old-world Zygonini tr. nov. (Hemiptera, Cercopoidea, Clastopteridae), with new taxa from the related Machaerotinae, Zootaxa 3768 (4), pp. 437-459 : 447

Ambonga apicenotata Melichar , redescribed

Ambonga apicenotata Melichar, 1915: 16 .

Diagnosis. Strongly hump-backed; tegmen strongly ampliated, 1.5 × as long as broad. Tawny, tegminal tip hyaline, marked with prominent brown patch curving from just beyond tip of clavus to nearly reaching costa; tegmen shiny, punctured from plaque to centre of clavus, surrounded by paler area in membrane; tip of tegmen edged with pale brown with veins weakly indicated and a prominent bulla below fork of Cu just before hyaline band across anteapical cells. Tylus broadly rounded, crown two-thirds as long as wide; head much narrower (0.60 ×) than pronotum; antenna with two basiconic sensilla just below a pair of coeloconic sensilla, and a single small pit beside this grouping; it also bears a prominent sensillum much closer to the ventral edge of the postpedicel ( Fig. 29 C). Tegmina with venation obscure, apical cells carinate, descending in size as in Pseudomachaerota grandis ( Maa 1962, fig. 6; see remarks under Hemizygon gen. nov.), but with inner 2 nearly square; hind wing as in P. grandis (idem, fig. 7H) but with 3 hooks on costal lobe, and cubital fork much broader, extending basad of 2nd apical cell. Ovipositor 1.4 mm long, tapered on apical third to a narrow point, with a very low tooth on dorsal margin just beyond midlength (as in Fig. 7 J). Male unknown. Length: female, 6.7 mm.

Type. Syntype female, MADAGASCAR: Mont. D’Ambre., Mar.; the only known type-specimen, No. 5192 in MMOE.

Remarks. The unique antennal sensilla suggest that coeloconic sensilla can be transformed into supernumerary basiconic sensilla, as appears to happen in some Aphrophorinae. However, these are not just abnormally formed sensilla, as their arrangement on the postpedicel also differs significantly from those of the sister-species, described below.

Figures

FIGURES 18 – 32. Antennae of Clastopteridae: Clastopterinae (18 – 20) and Machaerotinae: Hindolini (21 – 23), Machaerotini (24 – 25) and Zygonini (26 – 32, boxed set). A, lateral aspect of head; B, detail of antennal and antenna ledge; C, tip of antenna, showing postpedicel; D, detail of postpedicel showing detail of pits, revealing 4 enclosed coeloconic sensilla in communal pits. 18, Clastoptera; 19, Sepullia; 20, Grellaphia; 21, Chaetophyes; 22, Hindola; 23, Allox; 24, Blastocaena; 25, Irridiculum; 26, Pseudomachaerota cucullata; 27, Zygon, with detail showing basiconic sensilla (b) and coeloconic sensilla (c) of various types (shown in greater detail in Fig. 38); 28, Ambonga lanceolata (Zygonini); 29, Ambonga apicenotata syntype with insert showing sensilla at base of arista; 30, Pseudoclastoptera irrubesco sp. nov., holotype; 31 – 32, Pseudoclastoptera invidia sp. nov., male holotype (32) and unassociated female (31).

FIGURES 5 – 9. Zygonini, dorsolateral aspect (A); male subgenital plates (B), ventral aspect; (C), genital capsule and theca (with style in 7 – 9), lateral aspect; (D), theca, caudal aspect; (E) habitus, dorsal aspect. 5, Pseudoclastoptera irrubesco (style as in Pseudomachaerota); 6, Pseudomachaerota olivacea, female holotype; 7, Pseudomachaerota cucullata; 8, Hemizygon saturnus; 9, Zygon desegregatum. Scale line for habitus drawings: 1 mm.