Physocrotaphini, Chaudoir, 1863

Sabu, Thomas K., Abhitha, Prabhakar & Zhao, Danyang, 2008, A new Helluodes Westwood species from Western Ghats with notes on habitat, distribution and phylogenetic relationships of genera Helluodes Westwood and Physocrotaphus Parry of the tribe Physocrotaphini Chaudoir (Insecta: Coleoptera: Carabidae: Anthiinae), Zootaxa 1745 (1), pp. 30-46 : 32-33

Key to the genera of the tribe Physocrotaphini

1. Ligula prominent and distinctly longer than paraglossae ( Fig. 2 ); post-orbital surface area (gena) densely setose, with more than 40 setae and without tubercles; maxillary palpomere 4 shorter than 3; TLA = 24.5– 27.5 ( Fig. 23 ) ............................................................................................................. Helluodes Westwood.

- Ligula truncate and distinctly shorter than paraglossae ( Figs. 3–5 ); post-orbital surface area (gena) sparsely setose with less than 10 setae and with or without tubercles; maxillary palpomere 4 longer than 3; TLA = 19.5 ................................................................................................................................................ 2

2. Mentum with bifid median tooth; ligula stout and of uniform width from base to apex ( Fig. 3 ); male sternite 3 ( Fig. 15 ) with single oval depression bearing filamentous elongated setae; TLA = 19–19.5 ( Fig. 26 ) ................................................................................................................................... Physocrotaphus Parry.

- Mentum with triangular median tooth; ligula slender and gradually widening towards apex ( Figs. 4, 5 ); male sternite 3 with two oval depressions bearing filamentous elongated setae ( Fig. 16 )..........................3

3. Antennae longer, extended beyond the base of pronotum by 2–3 antennomeres; dorsal and ventral surfaces of antennomeres 5–11 devoid of impressions at base; antennomeres 2–11 with apical ring of elongate setae; mandibles long, narrow and weakly arcuate; anterior transverse sulcus of pronotum weakly developed; TLA = 7–17.6 ( Fig. 24 ) .................................................................... Pogonoglossus Chaudoir.

- Antennae shorter, extended beyond the base of pronotum by only the last; dorsal and ventral surfaces of antennomeres 5–11 bi-impressed at base; antennomeres 2–11 without apical ring of setae; mandibles short, wide and weakly arcuate; anterior transverse sulcus of pronotum very deep; length 14.5–16.8mm. ( Fig. 25 ) ............................................................................................................................. Schuelea Baehr.

Figures

FIGURES 2–5. Mentum (ventral view): 2 Helluodes devagiriensis n.sp.; 3 Physocrotaphus ceylonicus; 4 Pogonoglossus giganteus; 5 Schuelea drumonti. Scale bar = 1 mm.

FIGURES 23–26. Habitus: 23 Helluodes devagiriensis n.sp.(Holotype); 24 Pogonoglossus giganteus (Holotype); 25 Schuelea drumonti (Holotype); 26 Physocrotaphus ceylonicus (Lectotype). Scale bar = 5 mm.

FIGURES 15–18. Median depression and cluster of filamentous setae on 2nd and 3rd sternites of male: 15 Physocrotaphus ceylonicus; 16 Pogonoglossus giganteus; 17 Helluodes devagiriensis n.sp.; 18 2nd and 3rd sternites of female Helluodes devagiriensis n.sp. Not scaled.