Cylindronotum nevermanni (Liebke)

Erwin, Terry L., 2004, The beetle family Carabidae of Costa Rica: The genera of the Cryptobatida group of subtribe Agrina, tribe Lebiini, with new species and notes on their way of life (Insecta: Coleoptera), Zootaxa 662, pp. 1-54: 22-31

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Cylindronotum nevermanni (Liebke)


Cylindronotum nevermanni (Liebke) 

(Figs. 15, 16, 35)

Pseudometabletus nevermanni Liebke 1930: 723  . Holotype, male, COSTA RICA. LIMÓN, Rio Reventazon, Ebene, Hamburg Farm, 10 m, 10 ° 15 ' 0 N, 083° 28 ' 0 W, July ( NMNHAbout NMNH: ADP009555).

Cylindronotum nevermanni  van Emden, 1949: 891. New homonymy, new synonymy.

Derivation of specific name. The specific epithet, nevermanni  , was used by Liebke as a noun in apposition based on the last name of Ferdinand Nevermann, who was superb at collecting carabid beetles in Costa Rica, especially on his own farm (Hamburg Farm) and in its vicinity and who has contributed significantly to our knowledge of the species richness represented in Costa Rica. The NMNHAbout NMNH collection, where his collection was ultimately deposited, vouchers his efforts.

Common name. Nevermann's drab­brown carabid beetle.

Diagnosis. With the attributes of the genus as described above and color completely brown, except for testaceous 4 basal antennomeres; dorsal surface shiny.

Description. (Fig. 15). Size small: ABL = 4.66 to 5.08 mm, SBL = 4.33 to 4.66 mm, TW = 1.66 mm. Color: Brown to piceous overall. Luster: Surface shiny. Microsculpture: Effaced from head and pronotum; shallowly engraved isodiametric meshes on elytra. Head: Subequal across eyes to width of pronotum; frontal furrows absent except for small groove at margin of clypeus; frons slightly convex; eyes moderately small, slightly produced; ultimate labial palpomere sub­securiform; antennae of moderate length, reaching just posterior of humerus. Prothorax: Moderately convex, side margin narrowly beaded, evenly rounded at anterior marginal seta, moderately sinuate anterior to hind angle. Pterothorax: Normal for Agrina, fully winged. Legs: Normal for Agrina. Abdomen: Sterna normal for Agrina, setiferous, setae scattered, except normal ambulatory setae on sterna 3– 5 and both males and females with two pairs on sternum 6. Male genitalia: Phallus ( Fig. 16View FIGURE 16) with ostium of 1 / 3 its length, apex elongate, narrow; phallus narrow, twisted, catopic; internal sac with long flagellum, flagellum apex not barbed. Parameres asymmetric, left larger that right, right very small.

Dispersal potential. Adults are fully winged, thus they are likely good dispersers, as are most arboreal beetles.

Way of life. Beetles in this genus have been fogged from the canopy in South America. Wendy Porras, INBioAbout INBio parataxonomist, collected individuals by sweeping a plant in the family Convolvulaceae  . The known altitudinal range of this species is between 10 and 1300 meters above sea level. Adults have been collected only east of the Cordillera Central in March, June, July, and August, hence both dry and rainy seasons at low to middle altitudes.

Other specimens examined. COSTA RICA. 1 male, PUNTARENAS, 08° 29 ' N, 083° 36 ' W, August (D.R.Whitehead)( NMNHAbout NMNH: ADP059597); 1 female, Rincon, Golfito Dulce, Fundacíon Neotrópica Field Station, Quebrada Aquabuena below station, 75 m, 08° 42 ' 03” N, 083° 30 ' 49 ” W, June (W. Porras)( INBIO: ADP 108837); 1 female, 2.0 km S Santa Elena, 1300 m, 10 ° 18 ' 0 N, 084° 48 ' 0 W, June (J.M. & B.A. Campbell)( CNCAbout CNC: ADP057224); sex unknown, Estación Carara, R. B. Carara, 200 m, LN 195250,478700, March (R. Zuñiga)( INBIO: CRIAbout CRI 000164028). COLOMBIA. 1 female, META, Restrepo, December (J.A. Ramos)( NMNHAbout NMNH: ADP 100557). PANAMÁ. CANAL ZONE, Paraiso, 1 male, 1 female, February (E. Schwarz)( NMNHAbout NMNH: male, ADP08574, female ADP085572), 1 male, 3 females, March (E. Schwarz)( NMNHAbout NMNH: male, ADP08573, females ADP08571, 0 8567, 08566), Summit, 1 female, December (B. Bivin)( NMNHAbout NMNH: ADP 33323), Barro Colorado Island, 1 male, May (J. Zetek)( NMNHAbout NMNH: ADP08569), 1 male, Fort Sherman, January (H. Stockwell)( NMNHAbout NMNH: ADP 60083); PANAMÁ, 1 male, Panama City, April (A.H. Jennings)( NMNHAbout NMNH: ADP08570; CHIRIQUÍ, 3 males, 1 female, Bugaba, 244–457m, (G. Champion)( MCZAbout MCZ: female, ADP08563, male, ADP08560; NMNHAbout NMNH: males ADP08561, 08562).

Geographic distribution. ( Fig. 35View FIGURE 35). Known only from lower Central America from Costa Rica south to Colombia.

Notes. Cylindronotum nevermanni  van Emden was described from a female ( COSTA RICA. LIMÓN, Rio Reventazon, Ebene, Hamburg Farm, 10 m, 083° 28 'W, 10 ° 15 'N, September) by van Emden (1949), and supposedly deposited in the NMNHAbout NMNH (ibid: page 891). However, it is not in the NMNHAbout NMNH collection ( Erwin & House 1978). Apparently, van Emden was unaware that Liebke (1930) described the same species with the same patronymic and with the same Nevermann specimens, but in Liebke's own new genus placed in the wrong higher taxon, the Colliurinae.

), A. ignea Bates  (open circle,), A. lemoides Bates  (open box,), A. osa Erwin  n. sp. (diamond,), A. selva Erwin  n. sp. (closed star,), A. steineri Erwin  n. sp. (8 ­ point star,).

Figure 34. Geographical distribution map of the species of Cryptobatis  in Costa Rica: C. chontalensis Bates  (open box,





Smithsonian Institution, National Museum of Natural History


National Biodiversity Institute, Costa Rica


Canadian National Collection of Insects, Arachnids, and Nematodes


Universidade do Extremo Sul Catarinense, Bairro Universitário


Museum of Comparative Zoology














Cylindronotum nevermanni (Liebke)

Erwin, Terry L. 2004


Cylindronotum nevermanni

Emden 1949: 891


Pseudometabletus nevermanni

Liebke 1930: 723