Allorhynchium van der Vecht, 1963

Tan, Jiang-Li, Achterberg, Cornelis Van, He, Yi-Fei & Carpenter, James M., 2018, Northeast Asian Allorhynchium van der Vecht (Hymenoptera: Vespidae), with a key to Oriental species, Zootaxa 4434 (1), pp. 49-64 : 50-51

Allorhynchium van der Vecht, 1963

Figs 1–70

Allorhynchium van der Vecht, 1963: 57 –60; Baltazar 1966: 299; van der Vecht and Fischer 1972: 107; Cardale 1985: 165; Carpenter 1986: 64; Giordani Soika 1986a: 86, 1986b: 139–141, 1991: 163, 165, 1992: 41, 51, 1993: 29, 1996: 37; Gusenleitner 1988: 182, 1991: 262, 1997: 759, 1998: 508, 2000: 945; 2003: 855, 859, 2006a: 693, 2006b: 1304, 2007: 971, 2011: 1362; Nugroho et al., 2012: 75 –77; Nguyen et al. 2014: 8; Girish Kumar & Sharma 2014: 234, 238, 2015: 21–25; Pham & Li 2015: 134; Girish Kumar et al. 2016: 30 –34; Pannure et al. 2016: 22, 23. Type species (by original designation): Vespa argentata Fabricius, 1804 .

Archancistrocerus Giordani Soika, 1986b: 143 –146. Type species (by original designation): Archancistrocerus diffinis Giordani Soika, 1986 . Syn. nov.

Diagnosis. Vein 2r-m of fore wing closer to apex of marginal cell than to base of vein 1r-m, the fourth abscissa of the vein Rs usually only about half the length of the third ( Fig. 5 ); metanotum (“postscutellum”) at same level as mesoscutum or slightly oblique in lateral view ( Fig. 51 ), but never with pronounced declivous posterior part; propodeum medio-anteriorly near level of metanotum ( Fig. 51 ) or protruding ( Fig. 29 ); T1 often with tendency to form a transverse rim at the junction of the anterior and the dorsal faces (van der Vecht 1963) or having the junction of both faces angular in lateral view; sometimes S2 of ♂ armed with a flattened tubercle, two teeth or a transverse rim ( Fig. 4 ); S7 of ♂ with 1–3 teeth ( Fig. 64 ), pair of lobes or with a shiny and flat uplifted area ( Fig. 44 ); basal lamina of volsella with a long spine ( Fig. 60 ).

Notes. After examination of the holotype of Archancistrocerus diffinis Giordani Soika ( Figs 1–2 , 27–32 ) and the redescription of the male of Allorhynchium quadrimaculatum Gusenleitner ( Figs 4 , 56–65 ), it became obvious that the first species belongs to Allorhynchium van der Vecht. It has the typical distal position of vein 2r-m of the fore wing and a pronounced transverse (medially interrupted) rim on T1, the latter is also present in A. quadrimaculatum ( Fig. 59 ). This leaves only the vermiculate-rugose clypeus and the richer yellowish pattern of the body to separate Archancistrocerus from Allorhynchium ; both of these traits are considered too variable within Allorhynchium to be used as generic key characters.

The key below treats the Oriental species (including the southeastern Palaearctic area); both species restricted to Wallacea (Sulawesi and Moluccas) and Australia are excluded. Allorhynchium unifasciatum (von Schulthess, 1934) is reported by Giordani Soika (1986b) from China, but subsequently this species was designated as the genotype of Kennethia (Giordani Soika, 1994) , a junior homonym replaced by Orientalicesa Koçak & Kemal, 2010 . Additional key characters are given in square brackets.

Figures

FIGURES 1, 2. Allorhynchium diffinis (Giordani Soika), ♀, holotype. 2, Habitus, dorsal aspect; 3, ibid., lateral aspect.

FIGURES 3, 4. Allorhynchium spp., habitus, lateral aspect. 3, A. lugubrinum (Cameron, 1900), ♀, China; 4, A. quadrimaculatum Gusenleitner, ♂, Vietnam.

FIGURES 5–12. Allorhynchium argentatum (Fabricius), ♂, Indonesia (Sumatra). 5, wings; 6, head and mesosoma, lateral aspect; 7, mesosoma, dorsal aspect; 8, metasoma, lateral aspect; 9, head, anterior aspect; 10, head and pronotum, dorsal aspect; 11, antenna; 12, base of antenna.

FIGURES 13–19. Allorhynchium chinense (de Saussure), ♂, China (Henan). 13, head and mesosoma, lateral aspect; 14, propodeum and metasoma, dorsal aspect; 15, propodeum, latero-dorsal aspect; 16, terminal sternum (S7), lateral aspect; 17, head, anterior aspect; 18, head and pronotum, dorsal aspect; 19, propodeum, posterior aspect.

FIGURES 20–26. Allorhynchium chinense (de Saussure), ♀, China (Jiangxi). 20, wings; 21, head and mesosoma, lateral aspect; 22, propodeum and metasoma, dorsal aspect; 23, propodeum, latero-dorsal aspect; 24, propodeum, posterior aspect; 25, head, anterior aspect; 26, head and pronotum, dorsal aspect.

FIGURES 27–32. Allorhynchium diffinis (Giordani Soika), ♀, holotype. 27, wings; 28, head and mesosoma, lateral aspect; 29, mesosoma and first metasomal tergum, dorsal aspect; 30, metasoma, dorsal aspect; 31, head, anterior aspect; 32, head and pronotum, dorsal aspect.

FIGURES 33–40. Allorhynchium lugubrinum (Cameron, 1900), ♀. 33, right forewing; 34, head and part of mesosoma, lateral aspect; 35, ibid., dorsal aspect; 36, metasoma, lateral aspect; 37, ibid., dorsal aspect; 38, head, anterior aspect; 39, head and pronotum, dorsal aspect; 40, mesosoma, lateral aspect.

FIGURES 41–48. Allorhynchium metallicum (de Saussure), ♂, China (Tibet). 41, head and pronotum, lateral aspect; 42, propodeum and base of metasoma, dorsal aspect; 43, ibid., latero-dorsal aspect; 44, terminal sternum (S7), lateral view; 45, head, anterior aspect; 46, head and pronotum, dorsal aspect; 47, propodeum, posterior aspect; 48, apex of antenna.

FIGURES 49–55. Allorhynchium metallicum (de Saussure), ♀, China (Yunnan). 49, wings; 50, head and pronotum, lateral aspect; 51, propodeum and base of metasoma, latero-dorsal aspect; 52, ibid., dorsal aspect; 53, head, anterior aspect; 54, head and pronotum, dorsal aspect; 55, propodeum, caudal aspect.

FIGURES 56–65. Allorhynchium quadrimaculatum Gusenleitner, ♂, Vietnam. 56, wings; 57, head and mesosoma, lateral aspect; 58, mesosoma, dorsal aspect; 59, propodeum and metasoma, lateral aspect; 60, metasoma, lateral aspect; 61, antenna; 62, head, anterior aspect; 63, head and pronotum, dorsal aspect; 64, terminal sternum (S7), latero-ventral aspect; 65, apex of antenna.

FIGURES 66–71. 66, Allorhynchium concolor van der Vecht, ♀, paratype, Indonesia, head, anterior aspect. 67–69, A. snelleni (de Saussure), ♀ (66, Bawean) and ♂ (68, 69), ibid., but 69 detail of clypeus. 70, 71, A. vollenhoveni (de Saussure), ♀, Indonesia, ibid., but 61 detail of clypeus.