Hebrus syriacus Horváth, 1896,

Kment, Petr, Jindra, Zdeněk & Berchi, Gavril Marius, 2016, Review of West-Palaearctic Hebridae with description of a new species and redescription of Hebrus fulvinervis (Hemiptera: Heteroptera), Zootaxa 4147 (3), pp. 201-239: 217-219

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Hebrus syriacus Horváth, 1896


Hebrus syriacus Horváth, 1896 

( Figs. 26View FIGURES 23 – 28. 23 – 24, 29–39View FIGURES 29 – 35View FIGURES 36 – 39, 67View FIGURE 67)

Hebrus syriacus Horváth, 1896: 326  . Lectotype (designated by Linnavuori, 1994: 94): ♀, Israel, Haifa ( HNHM). 

Material examined. CYPRUS: Kyrenis, Ammos, 7.vii.1939, 1 ♀, H. Lindberg lgt. & det., P. Kment revid. ( NMPC). 

Descriptive notes. The female examined ( Figs. 26View FIGURES 23 – 28. 23 – 24, 29–39View FIGURES 29 – 35View FIGURES 36 – 39) fits the redescription by Linnavuori (1994), although the scutellum seems less bifid apically (see Figs. 36–37View FIGURES 36 – 39) and the membrane of the hemelytra is uniformly milky-brown and of the same colour as the posterior portion of the clavus ( Fig. 26View FIGURES 23 – 28. 23 – 24). The clavus has sparse pilosity and corium relatively dense pilosity ( Fig. 38View FIGURES 36 – 39). The membrane reaches the posterior 2/3 of mediotergite VII, leaving the apex of the dorsum free ( Fig. 26View FIGURES 23 – 28. 23 – 24).

Measurements (mm): Body length: 1.66. Head width (across eyes): 0.4. Pronotum: median length: 0.38, anterior width: 0.38, humeral width: 0.69. Lengths of antennal segments: I: 0.18, II: 0.14, III: 0.20, IVa: 0.08, IVb: 0.24. Length of metafemur: 0.51, metatibia: 0.63, metatarsus: 0.22.

Distribution ( Fig. 67View FIGURE 67). Asia: Cyprus (new record), Israel ( Horváth 1896, as Syria; Linnavuori 1961, 1994), Jordan (Linnavuori 1961),? Syria ( Stichel 1956, Andersen 1995). New species for Cyprus.

The type locality in the original description was given as ‘ Syria: Kaiffa (Reitter)’ ( Horváth 1896). Linnavuori (1961) listed four more specimens, one from Jordan and three from Judea, all from J. Sahlberg’s collection. Linnavuori (1994) interpreted the type locality as ‘ Israel, Haifa’ and speculated that ‘since all new finds from Israel belong to H. pusillus pusillus ( Fallén, 1807)  , H. syriacus  has possibly become extinct.’ Distribution of H. syriacus  in Syria requires confirmation as it seems to be based on the above mentioned error.

Comments. Horváth (1896) described the species based on the macropterous female, but did not specify the number of specimens, although there was probably only one. Andersen (1995) listed the type material as syntypes from Syria. Linnavuori (1994) redescribed the species based on a single female, presenting it as a ‘type’, and explicitely stated: ‘ The holotype is the only known specimen’. This statement fulfilled the requirements of Article 74.6 of the ICZN (1999) for fixation of the lectotype by inference of the holotype  .

Horváth (1896) compared the new species to H. ruficeps  : ‘Similar to H. ruficeps  , easily distinguished by body smaller and narrower, colouration of body pale, pronotum anteriorly not declivous, and posteriorly less dilated. Length 1.5 mm.’ Linnavuori (1994) redescribed the species based on the ‘holotype’ and provided illustrations of the head in lateral view and thorax and hemelytron in dorsal view, and provided the following differential diagnosis: ‘ H. syriacus  is readily distinguished from H. pusillus  and the related species by the smaller and narrower body, much narrower and more parallel-sided pronotum and, especially, the apically bifid scutellum’. Neither Horváth (1896) nor Linnavuori (1994) described apical antennal segments. Jordan (1954) included H. syriacus  in a comparative table of seven European Hebrus  species, and provided lengths of the antennal segments (mm): I— 0.18, II—0.16, III—0.20, IVa—0.10, IVb—0.24. However, the material Jordan (1954) used for comparison was not listed in that paper and remains unknown.

Linnavuori (1994) did not assign H. syriacus  to any of the subgenera; however, the metanotal elevation apically truncated with a wide V-shaped incision ( Linnavuori 1994: Fig. 4View FIGURES 4 – 12 b), antennal segments I and II subequal in length ( Stichel 1955, Linnavuori 1994), antennal segment IVb more than twice as long as IVa ( Jordan 1954), antennal segment I little longer than eye diameter ( Horváth 1896, Stichel 1955), and its small size (1.50–1.66 mm) suggest placement of H. syriacus  in Hebrusella  . (Body length of Euro-Mediterranean Hebrusella  is 1.30–1.80 (exceptionally 1.93) mm, in Hebrus  s. str. > 1.70 (usually 1.80) mm—see Poisson 1944, 1953; Jordan 1954; Wagner 1954, 1957; Kanyukova 1997, 2006)). On the other hand, examination of the antenna by scanning electron microscope revealed that segment IV is subdivided only by a membranous joint ( Figs. 34–35View FIGURES 29 – 35), which supports its placement in Hebrus  s. str. sensu Poisson (1944). However, the subgeneric classification of Hebrus  is problematic and requires revision (see Summary).


Hungarian Natural History Museum (Termeszettudomanyi Muzeum)


National Museum Prague














Hebrus syriacus Horváth, 1896

Kment, Petr, Jindra, Zdeněk & Berchi, Gavril Marius 2016

Hebrus syriacus Horváth, 1896 : 326

Linnavuori 1994: 94
Horvath 1896: 326