Sanaungulus longicornis, Yang & Zhao & Liu, 2024

Yang, Yuxia, Zhao, Wei & Liu, Haoyu, 2024, Taxonomic notes and new species of Burmomiles and Sanaungulus (Coleoptera, Cantharidae) from northern Myanmar during the late Mesozoic, Contributions to Zoology 2024 (2), pp. 153-177 : 161-164

Sanaungulus longicornis sp. nov.

(figs. 2, 3)

Etymology. The specific epithets derived from Latin “ longus ” (long) + “ cornus ” (antennae), referring to its long antennae, probably surpassing elytral apices if extending posteriorly.

Material. Currently only one specimen is known. This holotype is deposited in the Museum of Hebei University, Baoding, China ( MHBU, No. BU-COL-CAN0014) .

Horizon and locality. The lowermost Cenomanian (98.79 ± 0.62 Ma), mid-Cretaceous (Shi et al., 2012). Myanmar: Kachin State, Myitkyina District, Tanai Township, Hukawng Valley.

Differential diagnosis: The species looks similar to S. christensenae Fanti & Damgaard, 2019 and S. imparitibius Yang, Liu & Zhao, 2022 in antennomeres V–X each with a ramus, but can be distinguished from the latter two by the extremely long antennae, probably surpassing elytral apices if extending posteriorly, while at most extending to apical one-fourth in S. christensenae (Fanti & Damgaard, 2019: fig.7) and S. imparitibius (Yang et al., 2022: fig. 12).

Description. Male (figs. 2, 3). Total length 4.27 mm; width 0.71 mm. Body approximately 6.0 times longer than wide, dark brown.

Head (figs. 2, 3A) triangular. Eyes large and prominent, elliptical, inter-ocular distance approximately twice greater than the eye diameter. Mandibles invisible. Terminal maxillary palpomere approximately nearly as long as wide. Terminal labial palpomere twice as long as wide. Antennae (fig. 2) pectinate, probably surpassing elytral apices if extending posteriorly, scape stout and distinctly longer than pedicel, pedicel as long as III and expanding apically, V–X parallel-sided and each with an antennal appendage inserted at a basal cavity on the ventral side, XI bilaterally compressed and widened; relative length of antennomeres as 10:2:3:12:16:20:17:11:13:15:22.

Pronotum (figs. 2A, 3A–C) 0.51 mm long, 0.75 mm wide, nearly as wide as head, anterior margin arcuate, lateral margins parallel, posterior margin nearly straight, all margins bordered, anterior angles rounded, and posterior angles obtusely right-angled. Scutellum not well visible.

Elytra (figs. 2A, 3C) 2.27 mm long, 0.71 mm wide, at base nearly as wide as pronotum, approximately 3.2 times as long as width at humeri, lateral margins parallel and simple, apices rounded, surface finely punctate, punctures deep and rounded in striate, which distributed along with the moderately indicated longitudinal costae on disc. Hind wing slightly longer than elytra, not covering the abdomen completely.

Legs (fig. 2) long, meso- and metalegs distinctly longer than prolegs, femora stout, tibiae slender, pro-tibiae as long as femora, meso- and meta- tibiae slender and longer than femora, relative lengths of pro-, meso- and meta-tibiae as 10:13:19, tarsomere I as 10:13:20, and tarsomere II as 10:17:24.

Abdomen(figs.2A,3D,3E)slender.Terminal ventrite longer than terminal tergite, rounded at posterior margin. Terminal tergite widely troelsikloevedali is strongly transverse, with all and roundly emarginate in the middle of pos- margins nearly straight (Fanti & Damgaard terior margin, and acute at lateroapical angles. 2019: figs. 13, 14).

Aedeagus (figs. 3D, 3E) (only apical part vis- Description. Male (fig. 4A, B). Total length ible from the transparent tergite): lateral lobes 3.57 mm. Body yellow brown, elytra dark asymmetrical, separated and plate-like, ven- brown.

tral lobes larger than dorsal ones, right lobes Head (fig. 4A–C) 0.56 mm long, trianlarger than left ones, all lobes rounded at api- gular. Eyes large and prominent, elliptical. cal margins; median lobe enveloped by lateral Mandibles invisible. Terminal maxillary pallobes, enlarged at apex and roundly protrude pomere as long as wide. Terminal labial pal-

in the middle of posterior margin; laterophy- pomere twice as long as wide. Antennae (fig. ses well-developed, almost identical to each 4A–D) pectinate, probably surpassing elytral other, separated on both sides of median lobe, mid-length if extending posteriorly, scape rounded at apex. stout and longer than pedicel, III slightly Remarks. The yellow amber piece origi- longer than pedicel, IV distinctively longer nally measured 14 × 9 × 2 mm. The specimen than III, V–VIII parallel-sided and each with is complete. an antennal appendage inserted at a basal cavity on the ventral side, VIII–XI bilaterally compressed and widened, relative length of Sanaungulus elongaticollis sp. nov. antennomeres as 10:8:9:16:20:17:14:11:12:8:9.

(fig. 4) Pronotum (fig. 4A–C) 0.47 mm long, 0.26 mm wide, narrower than head, distinctly Etymology. The specific epithets derived from longer than wide, anterior margin arcuate, Latin “ elongatus ” (longer) and “ collum ” (neck), lateral margins rounded, posterior marreferring to its pronotum distinctively longer gin rounded, anterior and posterior angles than wide. rounded.

Material. Currently only one specimen Elytra (fig. 4B) 1.55 mm long, at base wider is known. This holotype is deposited in the than pronotum, abbreviated and uncover- Museum of Hebei University , Baoding , China ing the last four abdominal segments, lateral ( MHBU, No. BU-COL-CAN0015). margins parallel and simple, apices rounded, Horizon and locality. The lowermost surface finely punctate, punctures deep and Cenomanian (98.79 ± 0.62 Ma), mid-Creta- rounded in striate, which distributed along ceous (Shi et al., 2012). Myanmar: Kachin with the moderately indicated longitudinal State , Myitkyina District , Tanai Township , costae on disc. Hind wing slightly longer than Hukawng Valley . elytra, uncovering the abdomen completely. Differentialdiagnosis :Thespecieslookssim- Legs (fig. 4A, B) slender, all tibiae slender ilar to S. troelsikloevedali Fanti & Damgaard , and longer than femora, tarsomere I distinctly 2019 in the antennomeres V –VIII each with longer than II; relative lengths of pro-, meso- an appendage, triangular head and elliptical and meta-tibiae as 10:12:16, tarsomere I as eyes, but can be distinguished from the latter 10:17:26, and tarsomere II as 10:19:22 .

by the shape of pronotum, which is obviously Abdomen (fig. 4A, B, E) slender. Terminal longer than wide, with anterior margin arcu- ventrite longer than terminal tergite, rounded ate, lateral margins rounded, posterior mar- at posterior margin. Penultimate ventrite gin rounded. In contrast, the pronotum of S. slightly Downloaded emarginated from Brill in.com the 07 middle /10/2024of 02:00 the:56PM via Open Access. This is an open access article distributed under the terms of the CC BY 4.0 license. https://creativecommons.org/licenses/by/4.0/

posterior margin, and rounded at lateroapical angles. Terminal tergite rounded at posterior margin. Penultimate tergite rounded at posterior margin.

Remarks. The yellow amber piece originally measured 12 × 11 × 2.5 mm. The specimen is complete.