Wallabicoris dicrastyli, Schuh & P. Pedraza, 2010

Schuh, R. T. & P. Pedraza, 2010, Wallabicoris, New Genus (Hemiptera: Miridae: Phylinae: Phylini) From Australia, With The Description Of 37 New Species And An Analysis Of Host Associations, Bulletin of the American Museum of Natural History 2010 (338), pp. 1-118 : 31-34

Wallabicoris dicrastyli , new species Figures 8, 9; map 1; plate 1

DIAGNOSIS: Recognized by the chalkwhite coloration of the body and appendages (pl. 1), the moderately large size, with a mean

(AMNH_PBI 00132112).

total length of 5.41, and the structure of the male genitalia, the apical portion of the endosoma erect, the secondary strap of the endosoma complete and with a broad submedial undulation, and the left paramere very long, with an elevated shoulder on the posterior process, open over most of the dorsal surface, and greatly exceeding the margin of the pygophore in repose (figs. 8F, 9). Breeds on Dicrastylis flexuosa (Lamiaceae) . Smaller than the similarly colored Asteraceae-feeding W. baldersoni and W. schwartzi ; similar in size to the Asteraceaefeeding W. helichrysi and W. ozothamni ; larger than the Thymelaeaceae-feeding W. pinocchii and the Lamiaceae-feeding W. cassisi , W. lachnostachy os, W. newcastelii , W. paradicrastyli , and W. prostantheri . Potentially confused with W. rutidosi , but that species smaller, mean total length 4.87, and with a broken secondary endosomal strap.

DESCRIPTION: Male: Body weakly elongate, weakly ovoid, total length 5.22–5.64, ratio l/w 5 2.94. COLORATION (pl. 1): Pronotum unicolorous pale; hemelytron unicolorous pale, nearly white; markings on cuneus absent; membrane with elongate, contrasting, fumose marking at extreme base, remainder unicolorous pale, veins white; venter unicolorous pale or mostly so; antennal segment 1 unicolorous pale, with one or two black medial setae; antennal segment 2 unicolorous pale; labium pale with segment 4 heavily infuscate; hind femora with many brown or black spots; hind tibial spines dark with very small dark bases. SURFACE AND VESTITURE: Dorsum with reclining simple setae matching background coloration, and with some sericeous or woolly setae. STRUCTURE: Head: Somewhat projecting, ratio wh/lh 5 3.60 (fig. 8A); antennal fossa with ventral margin 1 diameter above ventral margin of eye; interocular space moderate, ratio iod/wh 5 0.44; eyes leaving gena broadly exposed in lateral view (hg5–14/ he20); labium reaching to midpoint of abdomen. GENITALIA (figs. 8E, F, 9): Endosoma: Base very long, with a tight Ushaped bend; distal half of shaft more or less straight and erect; primary endosomal strap weakly elongate apically, ratio lae/lsg 2.00; apex of primary endosomal strap nearly straight; secondary endosomal strap very slender, of uniform width from endosomal bend to gonopore, reaching to level of secondary gonopore, with a broad submedial undulation; secondary gonopore seen laterally in lateral view of endosoma. Phallotheca: More or less right angulate, dorsal surface with a hump; dorsal surface without a conspicuous keel. Left Paramere: Body very elongate, greatly exceeding margin of pygophore; in dorsal perspective open over nearly entire length; body tapered toward apex in lateral perspective; anterior process arising slightly forward of posterior margin of shaft; anterior process directed vertically; posterior process appearing tubular, fingerlike; posterior process with strongly projecting shoulder at base; base of posterior process at least somewhat elevated above level of paramere body. Right Paramere: Body moderately elongate, ratio lrp/wrp 3.68; body rounded laterally; posterior margin without distinct protuberance subapically; body more or less confluent with base; apex with a weakly elongate fingerlike process.

ETYMOLOGY: Named for the host genus, Dicrastylis J.L. Drumm. ex Harvey.

HOSTS: Recorded only from Dicrastylis flexuosa (W.R. Price) C.A. Gardner (Lamiaceae) (pl. 8B).

DISTRIBUTION (map 1): Known from the Goldfields region of Western Australia.

DISCUSSION: The known distribution of W. dicrastyli corresponds closely with the distributions of its known hosts in the Goldfields region of Western Australia.

HOLOTYPE: AUSTRALIA: Western Australia: 80.2 km W of Agnew toward Sandstone, 28.00117 ° S 119.9593 ° E, 650 m, 26 Oct 1996, Schuh and Cassis, Dicrastylis flexuosa (W.R. Price) C.A. Gardner (Chloanthaceae) , det. PERTH staff PERTH 05054729, 13 (AMNH_PBI 00135692) ( WAMP).

PARATYPES: AUSTRALIA: Western Australia: 80.2 km W of Agnew toward Sandstone, 28.00117 ° S 119.9593 ° E, 650 m, 26 Oct 1996, Schuh and Cassis, Dicrastylis flexuosa (W.R. Price) C.A. Gardner (Chloanthaceae) , det. PERTH staff PERTH 05054729, 173 (00090204, 00090208, 00090212, 00090221, 00090223, 00090225, 00090233, 00090555– 00090564), 21♀ (00090197–00090198, 00090200– 00090201–00090203,00090205–00090207, 00090209– 00090211, 00090215–00090220, 00090222, 00090224, 00090226) ( AM), 253 (00132109, 00132112, 00135676–00135679–00135681–00135691,00135693– 00135699), 46♀ (00135707, 00135713, 00135721– 00135764) ( AMNH), 63 (00090196, 00090199, 00090213–00090214, 00090227, 00090234), 7♀ (00090228–00090232,00090235–00090236)( UNSW), 43 (00132108, 00132110–00132111, 00132113), 32♀ (00090565–00090588, 00132121–00132122, 00135765–00135770) ( WAMP). NW Coastal Hiway 36 km N of Kalbarri Road, 27.62473 ° S 114.6902 ° E, 500 m, 29 Oct 1996, Schuh and Cassis, 13 (00373031) ( AM).

OTHER SPECIMENS EXAMINED: AUSTRA- LIA: Western Australia: 80.2 km W of Agnew toward Sandstone, 28.00117 ° S 119.9593 ° E, 650 m, 26 Oct 1996, Schuh and Cassis, Dicrastylis flexuosa (W.R. Price) C.A. Gardner (Chloanthaceae) , det. PERTH staff PERTH 05054729, 26♀ (00132114–00132120, 00135700–00135706, 00135708–00135712, 001- 35714–00135720) (AMNH). Carnarvon, Blue Bush, 24.88965 ° S 113.66922 ° E, 02 Sep 1953, A.S., 53 (00202579 [cat 57–327]–00202583 [cat 57–326]), 4♀ (00202575 [cat 57–316]–00202578 [cat 57–324]) (WAMP).