Wallabicoris craspedii, Schuh & P. Pedraza, 2010

Schuh, R. T. & P. Pedraza, 2010, Wallabicoris, New Genus (Hemiptera: Miridae: Phylinae: Phylini) From Australia, With The Description Of 37 New Species And An Analysis Of Host Associations, Bulletin of the American Museum of Natural History 2010 (338), pp. 1-118 : 28-30

Wallabicoris craspedii , new species Figure 6; map 3; plate 1

DIAGNOSIS: Recognized by the large size and the elongate slender body form, total length 5.56, ratio length/width 3.60, the heavy black setae on the dorsum, the more

(AMNH_PBI 00089293).

or less uniform greenish yellow coloration of the dorsum (pl. 1), and the form of the male genitalia, the secondary endosomal strap with a broad subbasal undulation, an elongate left paramere, and a lanceolate right paramere (fig. 6). Separated from other large species with a large elongate body as follows: from W. coolabah by the pale setae on the dorsum, the short submedial twist in the secondary endosomal strap, and the shorter left paramere in that species; from W. norsemanius by the pale setae on the dorsum, the longitudinal color pattern and the short submedial twist in the secondary endosomal strap in that species; and from W. uptoni by the less pronounced subbasal undulation in the secondary endosomal strap and the somewhat less elongate left paramere.

DESCRIPTION: Male: Body greatly elongate, parallel sided, total length 5.20, ratio l/w 5 3.63. COLORATION (pl. 1): Pronotum unicolorous greenish yellow; hemelytron mostly pale, greenish yellow; markings on cuneus present as partial or complete but weak infuscation; membrane with elongate, contrasting, fumose marking at extreme base, remainder weakly to strongly fumose over entire area, veins white; venter heavily infuscate, at least on mesopleuron and abdomen; antennal segment 1 dirty yellow, with strong infuscation at base and apex, with one or two black medial setae; antennal segment 2 dirty yellow proximally, weakly infuscate distally; labium generally infuscate, heavily so apically; hind femora with many brown or black spots; hind tibial spines dark with very small dark bases. SURFACE AND VESTITURE: Dorsum with erect black setae, particularly on pronotum, intermixed with pale setae on hemleytron, without sericeous or woolly setae. STRUCTURE: Head: Elongate, projecting anteriorly, ratio wh/lh 5 2.77; antennal fossa with ventral margin of fossa at ventral margin of eye; interocular space moderate, ratio iod/wh 5 0.42; eyes leaving gena moderately exposed in lateral view (hg3/he20); labium just reaching onto abdomen. GENITALIA (fig. 6): Endosoma: Base moderately long, with an open Ushaped bend; distal half of shaft more or less straight and erect; primary endosomal strap elongate apically, ratio lae/lsg 2.40; apex of primary endosomal strap nearly straight; secondary endosomal strap very slender, of uniform width from endosomal bend to gonopore, reaching to level of secondary gonopore, with a broad submedial undulation; secondary gonopore seen laterally in lateral view of endosoma. Phallotheca: More or less right angulate, dorsal surface with a hump; dorsal surface without a conspicuous keel. Left Paramere: Body elongate, somewhat exceeding pygophore margin; in dorsal perspective open over nearly entire length; body tapered toward apex in lateral perspective; anterior process arising at posterior margin of shaft; anterior process angled posterodorsally; posterior process appearing tubular, fingerlike; posterior process with strongly projecting shoulder at base; base of posterior process at least somewhat elevated above level of paramere body. Right Paramere: Body moderately elongate, ratio lrp/ wrp 3.38; body tapered from base to apex; posterior margin with a distinct protuberance subapically; body elevated at juncture with base; apex tapered.

ETYMOLOGY: Named for the host genus, Craspedia Forster (Asteraceae) .

HOSTS: Recorded from Craspedia sp. ( Asteraceae : Gnaphalieae ).

DISTRIBUTION (map 3): Known from several localities in New South Wales and Victoria, most in the Great Dividing Range.

DISCUSSION: Further attention to sampling on Craspedia will help to determine the extent to which this primarily high latitude— high elevation group of plants serves as hosts for Wallabicoris spp.

HOLOTYPE: AUSTRALIA: New South Wales: Mount Kosciusko, 36.45 ° S 148.26 ° E, 01 Feb 1952 – 12 Feb 1952, C.E. Chadwick, Craspedia sp. (Asteraceae) , 13 (AMNH_PBI 00089293) ( AM).

PARATYPES: AUSTRALIA: New South Wales: Charlottes Pass, Kosciusko National Park, 36.43333 ° S 148.31666 ° E, 1800 m, 17 Feb 1963, D.K. McAlpine, 13 (00090859) ( AM). Lake Cootapatamba, Snowy Mountains, 09 Feb 1979, D.K. McAlpine & B.J. Day, 13 (00090907) ( AM). Mount Kosciusko, 36.45 ° S 148.26 ° E, 01 Feb 1952 – 12 Feb 1952, C.E. Chadwick, Craspedia sp. (Asteraceae) , 2♀ (00089295, 00089296) ( AM); 14 Feb 1963, D.K. McAlpine, 13 (00090652) ( AMNH). Victoria: near Rosebud, Mornington Peninsula, 04 Jan 1976, M.S. Moulds, 13 (00090653) ( AM).