Claustropyga mirifica, Vilkamaa, Pekka & Hippa, Heikki, 2007

Vilkamaa, Pekka & Hippa, Heikki, 2007, Review of the Nearctic Claustropyga Hippa, Vilkamaa & Mohrig (Diptera, Sciaridae) with the description of three new species, Zootaxa 1552, pp. 53-68 : 60-62

Claustropyga mirifica sp. n.

Fig. 4 A–E

Material studied. Holotype male. CANADA, Quebec, Laniel, Cage # 54, 18.VIII.1933,?P. Ide ( CNC).

Male. Head almost unicolorous pale brown. Eye bridge 2–3 facets wide. Face (prefrons) with two setae. Clypeus non-setose. Antennal flagellomere 4, Fig. 4 A. Maxillary palpus, Fig. 4 B, with two palpomeres but elements of the two palpomeres weakly discernible in the apical one. Palpomere 1 with 1–3 setae. Palpomere 2 with 6–8 setae. Thorax dark brown, broadly paler brown on the middle part of pleura and laterally and at the row of dorsocentral setae on scutum. Scutum and scutellum as in C. brevichaeta ( Hippa et al. 2003, Fig. 3 a) but the number of lateral setae of scutum ca. 20, of which 4–5 are conspicuously stronger than the strongest dorsocentral setae; the number of scutellar setae ca. 16. Anterior pronotum with 1–2 setae. Episternum 1 with 3–4 setae. Other pleural sclerites non-setose. Wing, Fig. 4 C: pale yellowish-brown. Wing length 1.9 mm, width/length 0.50. c/ w 0.9. R1/R 0.7. Legs pale yellowish-brown, middle and hind coxa narrowly apically and their trochanters darker brown. Both fore legs lost in the specimen. Length of hind tibia/length of scutum and scutellum 1.8. Abdomen pale brown, concolorous with the paler areas of thorax. Sternite 8 with 8 setae, 5 on one side, 3 on the other, the latter may be damaged and this may be too low a number, all the setae in a transverse subapical row. Hypopygium, Fig. 4 D, E. Intercoxal lobe rather weak, with transverse posterior margin, without a distinct medial gap in setosity but the medial setae few in number. Gonostylus with a broad ventral mesial lobe with three closely-set megasetae. At the apex of gonostylus with two megasetae which almost completely coincide in the drawing. Apical tooth of gonostylus absent as well as megasetae additional to those mentioned above. Tegmen with a pair of apical finger-like lobes which are curved ventrad in their apical part. Setae of hypoproct (sternite 10) not observed in the specimen studied.

Discussion. Claustropyga mirifica is similar to the Palaearctic C. corticis but differs by the following characters: 1) The apical tooth of gonostylus is absent. 2) A strong megaseta on the mesial side of the gonostylus, in ventral view covered by the ventral mesial lobe, is absent. 3) The three megasetae at the margin of the ventral mesial lobe of the gonostylus are very close to one another so that in ventral view they touch or almost touch, whilst in C. corticis they are separated by caps which are as wide as or wider than the thickness of a megaseta. 4) There are two medially separate finger-like prongs apically on the tegmen, which are entirely absent in C. corticis . These finger-like prongs on the tegmen are quite unique in the Sciaridae .

Figures

FIGURE 3. Part of hypopygium (A, C) and gonostylus (B, D) in ventral view. A and B: Claustropyga auriculata Hippa, Vilkamaa & Mohrig (from Alaska). C and D: C. subcortici s Mohrig & Krivosheina (from Alaska). Scale 0.10 mm.

FIGURE 4. Claustropyga mirifica sp. n. (holotype): Antennal flagellomere 4 in ventral view (A), maxillary palpus in dorsal view (B), wing in dorsal view (C), part of hypopygium (D) and gonostylus (E) in ventral view. Scale 0.10 mm, in C 1.0 mm.