Scopocirinae

Ruiz, Gustavo R. S. & Maddison, Wayne P., 2015, The new Andean jumping spider genus Urupuyu and its placement within a revised classification of the Amycoida (Araneae: Salticidae), Zootaxa 4040 (3), pp. 251-279 : 273-274

Subfamily Scopocirinae Simon, 1901

Scopocireae Simon, 1901: 442, 1048 (Type genus: Scopocira Simon, 1900 ).

Monophyly: The two genera included here are only tentatively associated, as the DNA data for Gypogyna are weak, as are the morphological synapomorphies linking it with Scopocira . Males of both Scopocira and Gypogyna have apophyses on the distal prolateral portion of the chelicerae, but these are closer to the promarginal teeth, rather than on a more dorsal (external) position on the chelicerae. It is not clear whether these structures could be homologous to the amycine (plesiomorphic) mastidia or independently evolved projections. There is a dorsally placed RTA that appears to engage against a lobe of the cymbium, possibly to serve as a lock during copulation (see Costa & Ruiz 2014: figs 80–107). Both genera also share a feature unusual among spiders: males are typically larger than females. However, this is also seen, for example, in some gophoines.

Although Gypogyna is only tentatively placed with Scopocira , it can be placed with more confidence in the larger clade of node 2 ( Fig. 37 ) by molecular data, as well as by the form of the spermathecae and copulatory ducts, and the terminal embolus loop over the cymbium.

Figures

FIGURES 37 – 38. Maximum likelikhood phylogenies from Chimera and Non-chimera matrices, combined 5 gene analyses. Numbers show likelihood bootstrap % for Amycoids only; in parenthesis are bootstrap percentages for analysis without Nilakantha and Gypogyna (shown for deep amycoid branches only). Rectangles at nodes indicate bootstrap support in parsimony analysis (filled rectangle at least 75 %; open 50 % to 74 %). Sites partitioned as per PartitionFinder analysis.