Conescharellina angulopora ( Tenison Woods, 1880 )

Bock, Philip E. & Cook, Patricia L., 2004, A review of Australian Conescharellinidae (Bryozoa: Cheilostomata), Memoirs of Museum Victoria 61 (2), pp. 135-182 : 144

Conescharellina angulopora ( Tenison Woods, 1880)

Lunulites angulopora Tenison Woods, 1880: 7 , pl. 1 figs 3a–c.? Lunulites conica Haswell, 1881: 42 , pl. 3 figs 7, 8.? Conescharellina incisa Hincks, 1881: 127 (sep. p. 68), pl. 4 figs

1–3.? Bipora angulopora .— Whitelegge, 1887 ( 1888): 18. not? Lunulites angulopora .— MacGillivray, 1895: 46, pl. 8 fig. 1

(= Selenariopsis macgillivrayi Bock and Cook, 1996 ).? Conescharellina angulopora .— Levinsen, 1909: 311, pl. 23 figs

7a–f. not Conescharellina angulopora .— Gordon, 1985: 173, figs 20–23;

Gordon, 1989: 81, pl. 48B (see C. cognata ).

Remarks. Search for type material of Tenison Woods has been unsuccessful; consequently the characters of this species remain somewhat doubtful. The colony was figured as a distinct cone and the autozooids and avicularia occurred in apparent alternating radial series. However, the description of the orifice as “divided into two portions; one half triangular constricted in the middle; the other semicircular”, taken together with the illustration, indicates that Tenison Woods had confused the avicularia with the secondary orifices. The illustration shows at least one triangular avicularium accompanied by a typical lunate root pore that he did not recognise as distinct structures. His later remark “the cells are obliquely placed; sometimes in contrary directions alternately”, also appears to refer to avicularia, that have been described in other material assigned to this species as having alternating orientations. The description of “the vibracular pores” as “long and narrow, and in a depressed area” and the illustration, showing irregularly ovoid openings, apparently refers, in fact, to the secondary autozooid orifices. Waters (1887: 199), describing specimens he assigned to C. incisa (Hincks) , remarked “This may be Lunulites angulopora T. Woods , but apparently the avicularia were mistaken for zooecial cells, and the zooecia for vibracula”. Tenison Woods had only two specimens from Port Stephens, New South Wales, that he noted were “worn”; his type material has not been found. It seems unlikely that his species is recognisable. Livingstone (1924) regarded C. conica Haswell (1881) , Lunulites incisa Hincks (1881 , 1892), Bipora biarmata , and B. magniarmata Maplestone (1909) , all as junior synonyms of L. angulopora Tenison Woods (1880) . Both Livingstone (1928) and Hageman et al. (1996) reported C. angulopora from South Australia, and specimens labelled Bipora angulopora occur in Maplestone’s collection from this area. These specimens belong to at least two other taxa (see C. cognata and C. diffusa ) but specimens in Maplestone’s collection (NMV), inferred to have been from New South Wales, are described below as C. species ( C. angulopora sensu Maplestone not T. Woods). Haswell (1881) gave an illustration of his C. conica showing the orifices “upside down”, so that the apparent antapical primary sinuses are in fact, adapical parts of the peristome. He did not label his types or conserve entire specimens ( Livingstone, 1924). Hincks’ (1881) type specimens of L. incisa are not available, so that the identity of these species and their possible synonyms remains in doubt, in spite of the superficial similarity of his figure of L. incisa with that of C. conica (see below).