Bittacomorphinae, Alexander, 1920

Fasbender, Andrew & Courtney, Gregory W., 2017, A revision of Bittacomorphinae with a review of the monophyly of extant subfamilies of Ptychopteridae (Diptera), Zootaxa 4309 (1), pp. 1-69 : 22

publication ID

https://doi.org/10.11646/zootaxa.4309.1.1

publication LSID

lsid:zoobank.org:pub:C5C20BC9-B99C-47D0-970E-07E20D13CF72

DOI

https://doi.org/10.5281/zenodo.6027714

persistent identifier

https://treatment.plazi.org/id/03C18797-437C-C717-FF33-FF11FA5D0434

treatment provided by

Plazi

scientific name

Bittacomorphinae
status

 

Key to the extant species of Bittacomorphinae

Adults

This key was modified from that of Kang et al. (2012).

1. All legs with first tarsomere conspicuously expanded. Wing membrane without without macrotrichia. Antenna with 19–20 flagellomeres................................................................. Bittacomorpha Westwood ... 2

- All legs with first tarsomere not expanded. Wing membrane with or without macrotrichia. Antenna with 18–21 flagellomeres........................................................................... Bittacomorphella Alexander ... 3

2. Scutum dull black with silver-grey pollinosity laterally, prescutum with white stripe along medial suture. Cell m1 about onethird the length of cell r5 ( Fig. 7 View FIGURE 7 a). Male genitalia with apex of paramere weakly furcate ( Fig. 9 View FIGURE 9 f). Widespread east of Rocky Mountains................................................................ Bittacomorpha clavipes Fabricius

- Scutum shining black; pale medial stripe absent, but patch of silver-grey tomentum present on postsutural midline. Cell m1 about one-half the length of cell r5 ( Fig. 7 View FIGURE 7 b). Male genitalia with apex of paramere spatulate ( Fig. 10 View FIGURE 10 f). Coast and Cascade Ranges, northern Sierra Nevada .............................................. Bittacomorpha occidentalis Aldrich

3. Legs with three white bands............................................................................ 4

- Legs fewer than three white bands....................................................................... 5

4. Legs with 4th & 5th tarsomeres cream colored Japan............................... Bittacomorphella esakii Tokunaga

- Legs with 4th & 5th tarsomeres dark brown China.................. Bittacomorphella zhaotongensis Kang, Wang & Yang

5. Legs with narrow light band on proximal apex of 1 st tarsomere of foreleg................................................................................................ Taiwan and China ... Bittacomorphella lini Young & Fang

- Legs without light band on proximal apex of 1 st tarsomere of foreleg............................................ 6

6. Legs with two white bands............................................................................. 7

- Legs with one white band.............................................................................. 9

7. Legs with broad white band subapically on proximal portion of tibia. Male genitalia with posterior margin of epandrium platelike, posterior margin convex, epandrial claspers with basal lobe ( Fig. 16 View FIGURE 16 a). Eastern North America............................................................................................... Bittacomorphella jonesi Johnson

- Legs with narrow white band apically on proximal portion of tibia. Male genitalia with epandrium bilobate, posterior margin emarginate, epandrial claspers lacking basal lobe ( Fig. 17 View FIGURE 17 a)................................................... 8

8. Male genitalia with epandrial lobes semicircular. China............... Bittacomorphella gongshana Kang, Wang & Yang

- Male genitalia with epandrial lobes pentagonal ( Fig. 17 View FIGURE 17 a). Japan............... Bittacomorphella nipponensis Alexander

9. Wing without macrotrichia ( Fig. 7 View FIGURE 7 j). Thailand................................ Bittacomorphella thaiensis Alexander

- Wing with abundant macrotrichia ( Figs. 7 View FIGURE 7 c–e, g).......................................................... 10

10. Scutum with anterior of prescutum and presutural lateral portion of scutum shining dark brown/black, posterior apex of prescutum and medial portion of scutum with yellow stripe. Male genitalia with bilobate epandrium ( Fig. 15 View FIGURE 15 a). Gonostylus with a slender basal lobe directed dorsally ( Fig. 15 View FIGURE 15 b) Coast and Cascade Ranges of western North America. ........................................................................................ Bittacomorphella fenderiana Alexander

- Scutum black. Male genitalia with platelike epandrium ( Figs. 11 View FIGURE 11 a, 12a, 13a). Gonostylus without a basal lobe ( Figs. 11 View FIGURE 11 b, 12b, 13b)............................................................................................... 11

11. Scutum glabrous. Male terminalia with posterior margin of epandrium strongly emarginate, subaedeagal sclerite reduced, poorly distinguished plate at apex of aedeagus ( Figs. 11 View FIGURE 11 d, e), paramere bifurcate ( Fig. 11 View FIGURE 11 f) Coast Range of western North America. ................................................................. Bittacomorphella furcata sp. nov.

- Scutum covered in silver-grey tomentum. Male terminalia with posterior margin of epandrium weakly emarginate, subaedeagal sclerite of aedeagus conical, extending beyond phallotrema ( Figs. 12 View FIGURE 12 d–e, 13d), paramere simple. ( Figs. 12 View FIGURE 12 f, 13e).... 12

12. Male terminalia with subaedeagal sclerite of aedeagus tapering, apex with minute triangular tooth dorsally ( Fig. 12 View FIGURE 12 d, e). Paramere simple or with weak sigmoidal curvature ( Fig. 12 View FIGURE 12 f) Coast and Cascade Ranges of western North America. ............................................................................. Bittacomorphella pacifica Alexander

- Male terminalia with subaedeagal sclerite of aedeagus pin-like, apex simple ( Fig. 13 View FIGURE 13 d). Paramere forming semicircular curve dorsally ( Fig. 13 View FIGURE 13 e) Sierra Nevada ........................................... Bittacomorphella sackenii von Röder

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Diptera

Family

Ptychopteridae