Leptomorphus ascutellatus, Papp & Sevcik, 2011

Papp, L & Sevcik, J, 2011, Eight New Oriental And Australasian Species Of Leptomorphus (Diptera: Mycetophilidae), Acta Zoologica Academiae Scientiarum Hungaricae 57 (2), pp. 139-159 : 141-146

publication ID	https://doi.org/ 10.5281/zenodo.12612354
persistent identifier	https://treatment.plazi.org/id/03C38795-BE7D-FFF4-2625-4440EF2100F5
treatment provided by	Felipe
scientific name	Leptomorphus ascutellatus
status	sp. nov.

Treatment

Leptomorphus ascutellatus View in CoL sp. n.

( Figs 7–17 View Figs 7–13 View Figs 14–17 , 58–59 View Fig View Fig )

Holotype male ( HNHM): Thailand: Trang Prov., Thung Khai Botanic Garden , primary lowland rainforest, along the “Nature Trail”, Nov 13, 2004, No. 29, leg. L. PAPP & M. FÖLDVÁRI. (right 10 apical flagellomeres lost, some tarsomeres, incl. tarsomeres 4 and 5 lost, last tarsomere with claws remained on the right hind leg only)

Paratype female ( HNHM): same data .

Measurements: Body length 5.20 mm, head + thorax 1.23 mm, abdomen 3.97 mm (holotype),

c. 5.8 mm, 1.48 mm, c. 4.3 mm (abdomen strongly downcurved) (paratype female), respectively. Male. Body mostly metallic dark bluish grey. Head yellow, incl. palpi, scape and pedicel. Ocellar triangle black, ocelli comparatively large.

Anterior ocellus only slightly anterior to lateral ocelli. Distance of lateral ocelli only 0.08 m, head

0.55 and 0.60 mm broad, i.e. lateral ocelli far from eye margins. Postfrons shiny. Prefrons ventrally with 2 strong setae only. Eyes bare, dark grey, subshiny. Terminal palpomere 0.13 mm, ca. 6 times longer than broad and 1 1/3 times as long as penultimate one. Antennae dark grey. First flagellomere 0.115 × 0.093 mm, 2nd 0.21 × 0.01 mm, 7th 0.10 × 0.093 mm 14th (terminal) 0.125 × 0.07 mm. Terminal flagellomere not sharp apically but with a distinct central apex. Flagellomeres slightly flattened (strongly so in some related spp.).

pregenital (8th) tergite, perpendicular view. Scale bar: 0.4 mm for Fig. 2 View Figs 1–4 , 0.2 mm for Figs 1, 3–4 View Figs 1–4

Whole thorax dark, but scutellum yellow. Pleura around anterior spiracle dark. Scutellum very small ( Fig. 58 View Fig ) and so thinly sclerotised, that it may turn in to become slightly concave (as it is the case with the paratype female, Fig. 59 View Fig ). Scutellum entirely bare, not even microsetae present.

caudal view. Scale bar: 0.2 mm for Figs 7–9, 11 View Figs 7–13 , 0.1 mm for Figs 10, 12–13 View Figs 7–13

Legs yellow but base and apex of hind femur as well as base of hind tibia infuscated. Fore tibia 1.48 mm, fore basitarsus 2.11 mm long. Hind coxa 0.77 mm long (holotype). Microsetae on tibiae unaligned, rather short macrosetae present on all tibiae. Empodia not developed, claws minute (c. 0.03 mm).

Wing greyish with macrotrichia, veins ochre. All veins, incl. Sc, Rs and R-M setose. Sc ending in costa. Crossvein Sc-R present, close to apex of Sc. Length of R-M 0.20 mm, stem of medial fork 0.69 mm, M 1 1.38 mm, i.e. twice longer than M 1. Cu 2 close to, and only slightly divergent from Cu 1. Apex subdorsal) view. Scale bar: 0.2 mm for Fig. 14 View Figs 14–17 , 0.1 mm for Figs 15–17 View Figs 14–17

of Cu 2 overruns distinctly the level of Rs. M 3 c. 1.13 mm on holotype (wing broken there, so not precisely measurable), Cu 1 0.77 mm, M 3 of paratype female 1.54 mm, terminal section of Cu 1 0.80 mm, distance of their apices 0.86 mm. A 1 only slightly shorter than Cu 2, apex blunt. Halteres black.

Abdominal tergites 3 to 5 dark, each only with a short light band cranially.

Male terminalia: Setae asymmetrically placed on sternite 8 ( Fig. 7 View Figs 7–13 ). Tergite 8 larger, subspherical ( Fig. 13 View Figs 7–13 ). Genitalia very small, yellow. Cercal setae somewhat shorter than setae on hypoproct ( Fig. 13 View Figs 7–13 ). Gonocoxites higher than long or broad ( Figs 9, 11 View Figs 7–13 ), apical lobe of gonocoxite broad, slightly serrate caudally, more strongly apically ( Figs 10, 12 View Figs 7–13 ). Gonocoxites mostly fused with several long setae ( Fig. 11 View Figs 7–13 ), gonostyli mostly covered by gonocoxal lobes in caudal view.

Gonocoxites with an additional dark bilobed process at base of apical lobe, and those processes are perpendicular to the outer wall ( Fig. 12 View Figs 7–13 ). Gonostyli comparatively small but of an intricate structure ( Figs 12 View Figs 7–13 , 17 View Figs 14–17 ) with two extremely large black apical processes (we call them processes rather than thorns). Phallic organ ( Figs 15–16 View Figs 14–17 ) comparatively large. If we follow SØLI’ s (1997) interpretation (see SØLI 1997: fig. 30C), the paired, cranially directed structures ( Fig. 15 View Figs 14–17 ) must be named as parameral apodemes. Aedeagus largely triangular in dorsal (ventral) view. Aedeagal apodeme (a paired structure, too), better seen in lateral view, small ( Fig. 16 View Figs 14–17 ), ejaculatory apodeme large.

Female as male but postabdomen beginning with 8th segment yellow, 8th sternite with a patch of stiff black setae of c. 0.09 mm. Apical cercomere 0.23 × 0.165 mm.

Etymology. The specific epithet refers to the remarkably reduced scutellum.