Megalochus Khalaim & Broad

Khalaim, Andrey I. & Broad, Gavin R., 2013, Tersilochinae (Hymenoptera: Ichneumonidae) of Costa Rica, part 2. Genera Megalochus gen. nov. and Stethantyx Townes, Zootaxa 3693 (2), pp. 221-266 : 225-227

publication ID

https://doi.org/ 10.11646/zootaxa.3693.2.8

publication LSID

lsid:zoobank.org:pub:72B60375-3DF1-4EB1-B15E-587FDB6206BF

DOI

https://doi.org/10.5281/zenodo.6164300

persistent identifier

https://treatment.plazi.org/id/03C387F3-3A1E-7D7D-FBF6-E082FB80FEAB

treatment provided by

Plazi

scientific name

Megalochus Khalaim & Broad
status

gen. nov.

Megalochus Khalaim & Broad , gen. nov.

Type species: Megalochus grandis Khalaim et Broad , sp. nov.

Comparison. The new genus is related to Stethantyx , but differs from this genus by the slenderer first metasomal segment, lacking glymmae ( Fig. 21 View FIGURES 20 – 23 ), shortened antennae ( Fig. 20 View FIGURES 20 – 23 ), and propodeum and metapleuron with coarse irregular rugae ( Fig. 23 View FIGURES 20 – 23 ). These two genera form a distinctive group of genera, which we call the Stethantyx genusgroup, an addition to the three groups, Diaparsis , Phradis and Tersilochus genus-groups, suggested for European tersilochines by Horstmann (1981).

The Stethantyx genus-group differs from the other tersilochine genera by the fore wing with veins Rs +2 r and Rs angled more than 90° ( Figs 1–10 View FIGURES 1 – 6 View FIGURES 7 – 10 ), vein 2 rs-m and abscissa of M between 2 rs-m and 2 m-cu not or rather weakly thickened, and hind wing with vein cu 1& cu-a more or less vertical (usually reclivous in other tersilochine genera), and predominantly larger body size (4.2–15.0 mm). The wing venation of both genera seems plesiomorphic with respect to the usual tersilochine wing venation, with the unmodified veins 2 rs-m and M, the obtuse angled veins Rs +2 r and Rs and the narrow pterostigma. Megalochus is particularly un-derived, with hind wing vein M+Cu almost straight (strongly curved in all other tersilochine genera) and entirely pigmented, although not tubular basally (largely unpigmented, basally absent in other genera); the sub-basal cell is thus long and narrow, uniquely in the Tersilochinae . The wing venation, together with the short flagellomeres and narrowed proboscidial fossa are similar to the character states found in many Brachyscleromatinae, which may be closely related (cf. Quicke et al. 2009). It is tempting to speculate that Megalochus is a basal tersilochine that may partly bridge the gap between these two subfamilies, although this requires a phylogenetic test.

Diagnosis. Large species with body length 10.5–15.0 mm. Head strongly rounded behind eyes in dorsal view. Upper tooth of mandible longer than lower tooth. Clypeus large, lenticular, coarsely and densely punctate except lower margin. Malar space 0.5–0.6× as long as basal width of mandible. Flagellum of antenna unusually short, narrowed towards apex, with 27–36 flagellomeres ( Fig. 20 View FIGURES 20 – 23 ); all flagellomeres, except the basal and the apical ones, distinctly transverse. Occipital carina complete. Hypostomal carina absent. Mesoscutum flattened, low in lateral view ( Fig. 22 View FIGURES 20 – 23 ). Notaulus weak. Prepectal carina not joining to anterior margin of mesopleuron, dorsally indistinct ( Fig. 22 View FIGURES 20 – 23 ). Subtegular ridge strongly raised in frontal view, projecting laterally. Foveate groove of mesopleuron deep and long, coarsely crenulate, weakly curved, extending from anterior margin of mesopleuron to base of mid coxa ( Fig. 22 View FIGURES 20 – 23 ). Propodeum and metapleuron coarsely and irregularly rugose ( Fig. 23 View FIGURES 20 – 23 ). Distance between spiracle and pleural carina about half diameter of spiracle. Fore wing with vein 2 m-cu postfurcal. Veins Rs +2 r and Rs angled about 125°. Vein 2 rs-m twice longer than abscissa of M between 2 rs-m and 2 m-cu, both these veins not thickened.

Hind wing with vein cu 1& cu-a subvertical. Hind femur distinctly clavate ( Fig. 21 View FIGURES 20 – 23 ). Spurs of hind tibia weakly curved at apex. Tarsal claws not pectinate. Metasomal segment 1 slender, without glymma, round in cross-section ( Fig. 21 View FIGURES 20 – 23 ). Thyridial depression very long. Ovipositor upcurved, without teeth.

Etymology. From the Greece mega (large) and the latter part of the generic name Tersilochus , on account of the large body size. Gender masculine.

Composition. Only one species is known, M. grandis sp. nov., described below.

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