Dianous luteoguttatus, Champion, 1919

Tang, Liang, Wang, Wei, Szx, Olympus & Eos, Canon, 2018, Notes on the Dianous luteoguttatus complex with description of a new species (Coleoptera: Staphylinidae), Acta Entomologica Musei Nationalis Pragae 58 (1), pp. 237-242 : 242

Key to the Dianous luteoguttatus complex

1 Elytra without orange spots, metatarsomere I distinctly longer than the following three tarsomeres combined. ................................................................................ 2

– Elytra with orange spots, metatarsomere I distinctly shorter than the following three tarsomeres combined. ................................................................ 3

2 Punctation of interocular area dense, elytra longer with EL/EW=1.22–1.24. BL: 7.5–7.9 mm. Habitus: Fig. 5 ; sexual characters: Figs 15–17 . China (Yunnan). ........ ..................................... D. vorticipennis Puthz, 2005

– Punctation of interocular area sparse, elytra shorter with EL/EW=1.09–1.15. BL: 6.3–6.6 mm. Habitus: Fig. 6 ; sexual characters: Figs 18–22 . China (Sichuan, Yunnan). .................................... D. lilizheni sp. nov.

3 Femora bicolored. BL: 6.1–7.2 mm. Habitus: Fig. 2 ; sexual characters: Figs 7–9 . China (Yunnan, Xizang), Myanmar. .................. D. cruentatus L. Benick, 1942

– Femora uniformly black. ......................................... 4

4 Smaller species with BL: 5.3–6.1 mm, median portion of interocular area with interstices about as wide as diameter of punctures, punctation of elytra less confluent. Habitus: Fig. 4 ; sexual characters: Figs 11–14 . China (Guangxi, Fujian), Vietnam. ................ ..................................... D. depressifrons Puthz, 2016

– Larger species with BL: 6.0– 7.2 mm, median portion of interocular area with interstices distinctly narrower than half the diameter of punctures, punctation of elytra more confluent. .............................................. 5

5 Body with plumbeous luster, pronotal punctation less confluent, elytral spots smaller. BL: 6.0– 7.2 mm. Habitus: Fig. 1 ; sexual characters: Figs. 87, 90 in PUTHZ (2000), Fig. 1 in PUTHZ (1980). China (Taiwan), India, Nepal, Vietnam. ................................................ .............................. D. luteoguttatus Champion, 1919

– Body without plumbeous luster, pronotal punctation more confluent, elytral spots larger. BL: 6.0–7.0 mm. Habitus: Fig. 3 ; sexual characters: Fig. 10 ; see also Figs. 3, 6 in PUTHZ (2005). China (Yunnan). .............. ..................................... D. rimosipennis Puthz, 2005

Figures

Figs 1–6. Habitus of the species of the Dianous luteoguttatus species complex. 1 – D. luteoguttatus Champion, 1919; 2 – D. cruentatus L. Benick, 1942; 3 – D. rimosipennis Puthz, 2005; 4 –D. depressifrons Puthz, 2016; 5 –D. vorticipennis Puthz, 2005; 6 – D. lilizheni sp. nov. Scale bars: 1 mm.

Figs 15–22. Male sexual characters. 15–17 – Dianous vorticipennis Puthz, 2005; 18–22 – D. lilizheni sp. nov.; 15, 20, 21 – aedeagus; 16, 22 – sternite IX; 17, 19 – sternite VIII; 18 – tergites IX and X. Scale bars: 0.25 mm.

Figs 7–14. Sexual characters. 7–9 – Dianous cruentatus L. Benick, 1942; 10 – D. rimosipennis Puthz, 2005; 11–14 – D. depressifrons Puthz, 2016; 7, 11 – aedeagus; 8, 12 – male sternite IX; 9, 13 – male sternite VIII; 10, 14 – valvifer. Scale bars: 0.25 mm.