Cicurina (Cicurella) madla Gertsch 1992

Paquin, Pierre & Dupérré, Nadine, 2009, of 60 troglobitic species of the subgenus Cicurella (Araneae: Dictynidae), and a first visual assessment of their distribution, Zootaxa 2002, pp. 1-67 : 28-29

Cicurina (Cicurella) madla Gertsch 1992

( Figs 50–51, 134–135)

Cicurina madla Gertsch 1992: 109 , figs 91–92 (description of female). — Jackman 1997: 162, 171; Cokendolpher 2004a: 16, 19–20, 24–25, 27–28, 34, 42–46, 52, 54, figs 7–8, 40–47; Paquin & Hedin 2004: 3239, 3243–3250, 3253–3254; Reddell & Cokendolpher 2004: 80.

HOLOTYPE female: “ Texas; Bexar County, Madla’s Cave, located 1 mile west of Adam Wilson’s Cave Jr. October 4, 1963 J. Reddell, D. McKenzie ”. Examined ( AMNH).

Additional material examined. Christmas Cave , 06.ix.1993, 1♀ [ TMM #34448 ] J. Reddell, M. Reyes ( TMM); Headquarters Cave (Camp Bullis) 20.x.1995, 1♀, J. Reddell, M. Reyes ( TMM); Helotes Blowhole, no date, 1♀ 4 juvs [ TMM #34447 ], A. Grubbs, K. Kinsley, K. White ( TMM); Hills and Dales Pit ( UTSA area), 28.x.2000,1 ♀ [ TMM #34457 ] K. White, H. Becthol ( TMM); Lost Pothole (Government Canyon State Park), 04.ii.1995, 1♀, [ TMM #34449 ], A. Grubbs, G. Hoese, C. Vreeland ( TMM); Madla’s Drop Cave ,

Description. Female holotype

Total length: 7.29; carapace length: 2.70; carapace width: 1.67; opisthosoma length: 2.70. Eyes absent. Cheliceral promargin right side with 3 teeth (2 large, 1 small), left side likewise; retromargin right side with 6 denticles (2 large, 4 small), left side likewise. Legs I and IV missing. Epigynum: 0.448. Epigynal ventral plate with short transverse slit, bearing medial, very shallow indentation; atrium squared, two-thirds intruded by dorsal plate, atrial aperture small; bursa not enlarged nor fused; copulatory ducts touching at midpoint, reaching slightly over of primary lobe apex; copulatory ducts constriction located basally; primary pores not visible; stalk short, procurved, joining the primary lobe at the aperture of the dictynoid pore; dictynoid pore rounded; primary lobe elongated; secondary lobe absent; fertilization canal long, externally positioned from the dictynoid pore; fertilization duct medially attached to the primary lobe base.

Distribution. Reported from twenty caves in Bexar County, and one from Uvalde county. (Texas, Figs 134–135): Christmas Cave, Headquarters Cave, Helotes Blowhole, Hills and Dales Pit, Lost Pothole, Madla’s Cave (type locality), Madla’s Drop Cave, and Robber’s Cave. Paquin & Hedin (2004) provided localities for the species using codes. Here are the cave names that supported these records and the codes that were used: Fatman’s Nightmare Cave (NMF), Feature 50, UTSA (YFF), Helotes Hilltop Cave (LHH), Javelina Cave (CVJ), John Wagner Cave #3 (GWJ), La Cantera #1 (CAL), Lithic Ridge Cave (CRL), Margaritaville Cave (RAM) San Antonio Ranch Pit (RAS), Scenic Overlook Cave (LOS), Surprise Sink (SSC), Cranes Bat Cave (UEH, identity previously unknown). The Margaritaville Cave record from Uvalde County requires confirmation given the distance between the known distribution area and this locality (see figs 134–135).

Remarks. Male unknown. Data added by Gertsch (1992): “ 4 miles NW of Helotes” to facilitate location of the type locality. The occurrence of the species in some localities is based on a DNA match of immature specimens (see Paquin & Hedin 2004) and these specimens are not listed in the examined material. This species is included on the U.S. Federal list of endangered species ( Longacre 2000).