Epiphragma (Epiphragma) elongatum, Mao, Meng & Yang, Ding, 2009

Mao, Meng & Yang, Ding, 2009, New species of the genus Epiphragma Osten Sacken from Yunnan, Southwest China (Diptera, Limoniidae), Zootaxa 2121, pp. 44-56 : 53-55

Epiphragma (Epiphragma) elongatum sp. nov.

( Figs. 5 , 30–35 )

Diagnosis. Pronotum pale yellow, with two brown stripes. Femora yellow with two dark brown rings, first at 2/3 length of femora, second at tip, broader. Cross-vein m-cu at near 1/4 length of cell dm. Outer gonostylus with base very broad and tip curved into spine. Inner gonostylus very deplanate with obtuse tip. Interbase with a very long and slender rod (nearly as long as gonocoxite) beyond base, apex a sharp point; basal half of base finger-shaped, apical half enlarged and rounded.

Description. Male. Body length 10.8 mm, wing 12.5 mm.

Head. Yellowish brown with pale gray pollen except orbit pale yellow. One median tubercle between eyes. Hairs on head black. Antenna 2.5 mm. Scape and pedicel brown, flagellum 13-segmented with two basal segments fused, fusion-segment yellow, succeeding segments brownish black, flagellomeres cylindrical, apical segments elongate, with longer verticils. Proboscis yellow with black hairs; palpus brownish black with black hairs.

Thorax. General brownish yellow with pale gray pollen. Pronotum pale yellow, with two brown stripes. Prescutum with four black stripes, intermediate pair long and almost extending to transverse suture. Lateral, humeral and cephalic parts light yellow. Scutum, scutellum and mediotergite yellowish brown. Pleura brownish yellow, variegated by darker areas. Hairs on thorax brown. Coxae and trochanters yellow; femora yellow with two dark brown rings, first at 2/3 length of femora, second at tip, broader; tibiae brown with one spur; tarsi brown. Hairs on legs black except coxae with brownish yellow hairs.

Wing ( Fig. 5 ) with conspicuous brown patterning, chiefly ocelliform, with narrow dark brown margining. Base of wing brown, connected with two large ocellate circles with flattened tips; one irregular broad band along cord connected with second circle; one M-shaped patterning around outer end of cell dm, connected with one irregular patterning along wing margin, leaving five pale spots at tips of cells R3, R4, R5, M1, M2; one dark area occupied both posterior half of cells Cu and A1, only with one pale spot at 3/4 length of cell Cu; one additional dark area in almost whole cell A2, with two small pale spots. Stigma solidly dark brown. R2+3+4 longer than R2+3; m-cu at near 1/4 length of cell dm. Halter 1.8 mm long, yellowish brown except base of stem light yellow and apex of knob pale yellow.

Abdomen. Tergites blackish brown, with one narrow median brownish yellow line, margined with brownish yellow on posterior half of each tergite. Sternites 1–7 brown, sternites 8–9 black. Hairs on abdomen brown.

Hypopygium ( Figs. 30–35 ). Posterior margin of tergite 9 ( Fig. 30 ) with subtrigonal lobe on each side of median V-shaped notch. Base of outer gonostylus ( Fig. 31 ) very broad, tip curved into spine. Inner gonostylus ( Fig. 32 ) very deplanate, with tip obtused. Interbase ( Fig. 34 ) with a long and very slender rod beyond base, nearly as long as gonocoxite, apex a sharp point; basal half of base finger-shaped, apical half enlarged and rounded.

Female. Body length 12.6 mm, wing 14.2 mm. Similar to male. Cerci reddish brown; hypovalves yellowish brown.

Type material. Holotype male, Yunnan: Tengchong, Zizhi villiage (1750 m), 25°43΄44ʺN 98°34΄0 7ʺE, 2007. V. 31, Xingyue Liu. Paratype 1 female, same data as holotype.

Distribution. China (Yunnan).

Etymology. The specific name refers to the elongated interbase.

Remarks. This new species is somewhat similar to E. rhododendri Alexander, 1966 from India (Sikkim) in having the similar patterning of the femora, but it can be separated from the latter by the flagellum 13- segmented with two basal segments fused, outer gonostylus very broad at base, inner gonostylus very deplanate with the obtuse tip, and interbase with a long and very slender rod (nearly as long as the gonocoxite) beyond the base. In E. rhododendri , the flagellum is 12-segmented with three basal segments fused; the outer gonostylus is relatively narrow; the inner gonostylus is longer, the apical half is a paddlelike blade, and the apical spine of the interbase is subequal to or slightly longer than the enlarged base, gradually narrowed to the acute tip ( Alexander, 1966).

Figures

FIGURES 1 – 5. Wing (male). 1. Epiphragma (Epiphragma) mediale sp. nov.; 2. Epiphragma (Epiphragma) yunnanense sp. nov.; 3. Epiphragma (Epiphragma) ancistrum sp. nov.; 4. Epiphragma (Epiphragma) breve sp. nov.; 5. Epiphragma (Epiphragma) elongatum sp. nov.

FIGURES 30 – 35. Epiphragma (Epiphragma) elongatum sp. nov. Male. 30. Ninth tergite, dorsal view; 31. outer gonostylus, lateral view; 32. inner gonostylus, inner lateral view; 33. hypopygium, lateral view; 34. hypopygium, dorsal view; 35. hypopygium, ventral view.