Colocasiomyia besaris Yafuso & Toda, 2023

Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak & Toda, Masanori J., 2023, Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species, Zootaxa 5278 (2), pp. 201-238 : 223-224

publication ID	https://doi.org/ 10.11646/zootaxa.5278.2.1
publication LSID	lsid:zoobank.org:pub:697AC989-141D-4A89-8F02-5A4644E303EA
DOI	https://doi.org/10.5281/zenodo.7907978
persistent identifier	https://treatment.plazi.org/id/03C687D9-FFC0-FFF7-A5C6-F9E5838AE1F9
treatment provided by	Plazi
scientific name	Colocasiomyia besaris Yafuso & Toda
status	sp. nov.

Treatment

2) Colocasiomyia besaris Yafuso & Toda View in CoL , sp. nov.

( Figs 2–12B View FIGURE 2 View FIGURE 3 View FIGURE 4 View FIGURE 5 View FIGURE 6 View FIGURE 7 View FIGURE 8 View FIGURE 9 View FIGURE 10 View FIGURE 12 , 14B View FIGURE 14 )

Colocasiomyia sp.3 aff. nepalensis: Sultana et al., 2006: 694 View in CoL .

Diagnosis. Surstylus with 8–10 stout, longer, peg-like teeth; dorsalmost one thickest (Fig. 11-1B 2). Pregonites apically pointed, smooth on ventral margin (Fig. 11-1B 4). Epigynium with 26–28 small sensilla per side near lower posterior margin ( Fig. 12B View FIGURE 12 1 View FIGURE 1 ). Distal, narrow portion of hypogynial valve longer than basal, broad portion ( Fig. 12B View FIGURE 12 ).

Description (♁ and ♀). Head. Antennal pedicel blackish brown; first flagellomere approximately 1.6 times as long as pedicel; arista approximately 2.1 times as long as first flagellomere ( Fig. 3B View FIGURE 3 ). Distance between antennal scapes as wide as scape diameter ( Fig. 2B 2 View FIGURE 2 ). Facial carina wider than 1st flagellomere ( Fig. 2B 2 View FIGURE 2 ). Supracervical setae 8–10 and postoculars 18–20 per side ( Fig. 4A View FIGURE 4 ). Cibarial projections at anterolateral corners as long as width of anterior margin; medial sensilla approximately 2–3 per side ( Fig. 5B View FIGURE 5 2 View FIGURE 2 ). Prementum and neighboring lateral membrane with 9–10 setae per side ( Fig. 5B View FIGURE 5 2 View FIGURE 2 ). Labellum with 11 pseudotracheae per side ( Fig. 5B View FIGURE 5 1 View FIGURE 1 ).

Thorax. Two katepisternal, prominent setae longer than acrostichal setulae ( Fig. 7B View FIGURE 7 ).

Legs grayish brown, except for all tarsi pale grayish yellow ( Fig. 14B View FIGURE 14 ). Fore tarsomere I slightly longer than tarsomeres II and III combined; mid I as long as II+III+IV; hind I slightly longer than II+III+IV ( Fig. 9B View FIGURE 9 ). Fore tarsomere II with 1 large and 1 small pegs; latter 1/3 as long as former ( Fig. 9B View FIGURE 9 1 View FIGURE 1 ).

Abdomen. Tergites blackish brown ( Fig. 14B View FIGURE 14 ). Male sternites III and IV as wide as long; V+VI distally deeply bilobed ( Fig. 10B View FIGURE 10 1 View FIGURE 1 ). Female sternites III–V as wide as long; VI wider than long; VII wider than long plate bilobed in distal half ( Fig. 10B View FIGURE 10 2 View FIGURE 2 ). Female tergite VII mid-dorsally separated into 2 lateral plates.

Male terminalia. Epandrium with 2–4 long setae only on lateral to ventral portion (Fig. 11-1B 1). Cercus nearly entirely pubescent except for roundish, ventral flap, with approximately 40 setae (Fig. 11-1B 1). Phallal sheath partly pubescent (Fig. 11-1B 4).

Female terminalia. Perineal membrane with patch of dense, distinct warts.

Measurements (holotype /range in 2♁ and 1♀ paratypes, in mm): BL = 1.51/♁ 1.35–1.55, ♀ 1.70; ThL = 0.67/♁ 0.72–0.73, ♀ 0.72; WL = 1.39/♁ 1.46–1.55, ♀ 1.47; WW = 0.66/♁ 0.72, ♀ 0.72.

Indices (holotype /range in 2♁ and 1♀, or less if noted, paratypes, in ratio): FW/HW = 0.54/0.53–0.56; ch/o = 0.38/0.32–0.40; prorb = 0.83/0.82–0.97; rcorb = 0.30/0.25–0.32; vb = 0.21/0.24–0.29; dcl = 0.51/0.52–0.54 (1♁ and 1♀); presctl = 0.30/0.35–0.41; sctl = 0.75/0.71–0.73 (1♁ and 1♀); sterno = 1.07/1.28–1.42; orbito = 0.35/0.38–0.45; dcp = 0.58/0.68–0.77; sctlp = 0.81/0.91–1.07; C = 2.10/1.82–2.18; 4c = 1.32/1.22–1.39; 4v = 2.54/2.33–2.61; 5x = 2.78/1.82–3.23; ac = 2.50/2.36–2.87; M = 0.99/0.91–1.02; C3F = 0.41/0.46–0.54.

Holotype. ♁ ( MZB), “ Cibodas , West Java, Indonesia, 12.xii.2004, K. T. Takano / ex inflorescence of Pinanga coronata ”.

Paratypes. Indonesia: 12♁, 4♀, same data as the holotype ( MZB, SEHU); 2♁ (#05200–01) , 1♀ (#05226), same data as the holotype except for 24.xii.2003 ( KIZ) .

Distribution. Indonesia (West Java).

Remarks. This species resembles C. pinangae sp. nov., but can be distinguished from it by the diagnostic characters.

Etymology. The specific name derives from “besar” (meaning “large”) of Indonesian, referring to its larger body size among the members of the zeylanica group.