Diplommatina exserta Godwin-Austen, 1886

Budha, Prem B., Naggs, Fred & Backeljau, Thierry, 2017, The genus Diplommatina Benson, 1849 (Gastropoda: Caenogastropoda: Diplommatinidae) in Nepal, with the description of seven new species, European Journal of Taxonomy 337, pp. 1-30 : 8

Diplommatina exserta Godwin-Austen, 1886

Figs 3B , 9B

Diplommatina exserta Godwin-Austen, 1886: 184 , pl. 49, figs 2, 2a.

Diplommatina (Diplommatina) exserta – Kobelt 1902: 432.

Diplommatina exserta – Gude 1921: 314. — Budha et al. 2015: 6.

Diagnosis

Measurements (n = 3): SH 3.4–3.5 mm, SW 1.7–1.8 mm, Wh 7.5. Dextral, apical whorls slowly increasing in size, resulting in a slender conical spire. Penultimate and body whorls nearly equal in width. Constriction between parietal lips of the peristome at AeP ( Fig. 9B ). Two parietalis: Prt1 long and strongly developed, gradually elevated inner side of the body whorl, Prt2 short ( Fig. 3B3 ). Two vertical palatalis: Vpt1 stronger and close to the constriction,Vpt2 short. One long horizontal palatalis, touching the inner parietal lip ( Fig. 9B ). Columellaris strong. Columellar tooth prominent. Protoconch smooth, Wh 2.0, with minute evenly distributed pits, no radial ribs ( Fig. 3B 1 ). Teleoconch with widely spaced radial ribs; 7–8 ribs/ 0.5 mm on the penultimate whorl; 8–10 ribs/ 0.5 mm on the body whorl towards the aperture ( Fig. 3B 2 , B 4 ). Ribs moderately prominent. Very fine spiral striation present. Umbilicus open. Aperture inversely C-shaped, due to the straight columellar edge of the lip. Peristome with two close lips; under high magnification 2–3 ribs seen between the two lips. Parietal wall of the aperture extending to nearly halfway up the body whorl.

Material examined

NEPAL: 4 shells, Siddha Cave, Tanahun, 695 m, 27.94544 N, 84.422244 E, 24 Oct. 2008, P.B. Budha and R. Devkota leg. ( CDZMTU 024).

Remarks

The most similar species are Diplommatina exilis W.T. Blanford, 1863 from Myanmar ( Hanley & Theobald 1876: pl. 119, fig.10) and D. pullula Benson, 1859 from “Rungun on the West of Darjiling” ( Hanley & Theobald 1876: pl. 119, fig.7). Compared to D. exserta , the apical whorls are wider in D. pullula and more tumid in D. exilis .

Figures

Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm.

Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov.

Fig.1. Map of the localities and areas surveyed for Diplommatina spp. in Nepal.

Fig. 2. Shell terminology. A. D. regularis Fulton, 1901. B. D. folliculus (L. Pfeiffer, 1846). C. General outline of the lower part of a shell from Diplommatina species. Abbreviations: Prt = Parietalis (Prt1 = inner parietalis, Prt2 = outer parietalis); Pt = Palatalis; Hpt = Horizontal palatalis (Hpt1 = lower, Hpt2 = upper in case of two); Vpt = Vertical palatalis (Vpt1 = left side, Vpt2 = right side in case of two).

Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm.