Bembidion cachagua, Maddison, 2023

Maddison, David R., 2023, A New Subgenus of Bembidion Latreille from México and Guatemala, with Descriptions of Two New Species (Coleoptera: Carabidae), Proceedings of the California Academy of Sciences 67 (19), pp. 433-450 : 446

Bembidion cachagua , sp. nov.

( Figures 7 , 8)

Holotype male, in CAS, labeled: “ U.S.A.: CALIFORNIA: Monterey Co., Hastings Nat. History Reserve. J.M. Linsdale Colln. ”, “ C.A.S. Accession 29-V-48 Cachagua Creek ”, “David R. Maddison DNA3842 DNA Voucher” [pale green paper], “ HOLOTYPE Bembidion cachagua David R. Maddison” [partly handwritten, on red paper]. Genitalia mounted in Euparal in between coverslips pinned with specimen; extracted DNA stored separately.

Paratypes (2 males, 4 females). Same data as holotype (3: CAS, OSAC) . “ Pinnacles Nat’l Mon. 4.19.58 Toschi ” (1: EMEC; this is D.R. Maddison DNA voucher number 4758, and EMEC specimen number 348099). “Santa Barbara Cal. May ’ 91 L. E.R.” (1: MCZ, Hayward Collection). “ S. Cal. ” (1: UIUC) .

Type locality. Cachagua Creek does not flow through Hastings Natural History Reserve , although a tributary, Finch Creek , does. At its nearest Cachagua Creek is approximately 2.7 km from the boundary of the Reserve. It is reasonable to presume that the upper label on the holotype, “ U.S.A.: CALIFORNIA: Monterey Co., Hastings Nat. History Reserve. J.M. Linsdale Colln. ”, was a generic label used for all specimens from the Linsdale collection, as J.M. Linsdale was the Director of the Hastings Natural History Reserve at the time ( Alagona, 2012). The lower label presumably allowed customization for each specimen, and the locality information on the second label is thus likely more accurate. Thus, the type locality would more accurately be stated to be “Cachagua Creek near Hastings Natural History Reserve” .

Derivation of specific epithet. Named after Cachagua Creek, the locality at which four of the seven known specimens were found. Treated as a noun in apposition.

Description. Body length 3.45–3.90 mm. Forebody dark brown or piceous, with metallic greenish-yellow or brassy sheen; elytra pale chestnut brown, with the four testaceous elytral spots contrasting only slightly against the ground color ( Fig. 7 ). In four specimens, the antennae are infuscated throughout; in the other three specimens, two of which appear to be teneral, the basal 2.5 antennomeres are testaceous; legs pale reddish brown, slightly infuscated. Frontal furrows shallow, not prolonged onto clypeus. Mentum with anterior lateral regions triangular, large; central tooth of mentum triangular, slightly rounded. Prothorax narrow, only slightly wider than head; hind angles slightly obtuse; posterolateral carina of pronotum long and more or less straight; posterior region of pronotum rugose, wrinkled and with evident punctures ( Fig. 7 ). Elytra with lateral bead not prolonged medially at shoulder; striae shallow, poorly defined, with extremely small punctures. Microsculpture evident throughout the dorsal surface of the forebody in most specimens, more or less isodiametric or slightly transversely stretched, in some specimens effaced in the center of the head between the eyes, and in the center of the pronotal disc; microsculpture effaced from most of elytra, but evident behind the second dorsal discal seta (ed5) as defined, slightly transverse sculpticells. Pronotum with two lateral setae on each side; elytron with two setae in third stria. Aedea-

Figures

FIGURE 7. Holotype of Bembidion cachagua. (A) Habitus, scale bar 1 mm. (B) Forebody, scale bar 0.5 mm.