Pupennea Páll-Gergely, 2020

Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Sajan, Sheikh, Tripathy, Basudev & Chen, Zhe-Yu, 2020, A review of the Diapheridae (Gastropoda: Eupulmonata: Streptaxoidea), with special emphasis on India and Myanmar, Raffles Bulletin of Zoology 68, pp. 682-718 : 700

Pupennea Páll-Gergely , new genus

Type species. Pupa planguncula Benson, 1863 .

Diagnosis. As in the type species.

Differential diagnosis. For homologies of apertural denticles, see Table 1. Pupennea , new genus, differs from all Sinoennea species by the short parietal lamella, which is long and consists of two portions even in small-shelled members of Sinoennea . The columellar lamella is pointed, small, and situated in the parieto-columellar corner. In contrast, most Sinoennea species have a deeply-situated, blade-like columellar lamella. Anterior of the columellar lamella there is a strongly developed columellar denticle, which is not present in Sinoennea . Pupennea , new genus, possesses an inner and outer basal denticle. In a few Sinoennea species one basal denticle is present, which is probably homologous with one of the basal denticles of Pupennea , new genus. The parietal callus is only indicated in S. planguncula , but well-developed in most Sinoennea species.

Platylennea , new genus, is similar to Pupennea , new genus, in possessing both a columellar lamella and a columellar denticle. Furthermore, two basally situated teeth are found in both genera. However, the homologies of the latter structures are questionable (see Table 1), because in Platylennea the lower basal denticle is situated closer to the columella and to the peristome, whereas in Pupennea the tooth situated closer to the columella (inner basal denticle) is the one, which is situated deeper in position. Further differences between the two genera are in the shell shape (ovoid in Pupennea , new genus, and elongate conical in Platylennea , new genus), the much shorter parietal lamella in Pupennea , and the more complex palatal teeth.

Rowsonia , new genus, has a well-developed parietal lamella, lacks a columellar denticle, and in the position of the upper palatal tooth there is only a slight thickening (a complex bilobed tooth in case of Pupennea , new genus). Furthermore, the deeply situated palatal folds of Rowsonia are absent in Pupennea .

The strongly developed upper parietal tooth, corresponding with a depression on the outer side of the palatal wall suggests that S. planguncula is related to Sinoennea instead of Rowsonia , new genus, and Platylennea , new genus.

Included species. Pupennea planguncula ( Benson, 1863) .

Etymology. The name Pupennea is the combination of the generic names Pupa, which was often used to be used for ovoid shells in the 19 th century, and Ennea H. Adams & A. Adams, 1855 , which is a generic name that used to be widely used for high-spired Streptaxidae . Grammatical gender: feminine.

Distribution. Central India ( Fig. 10 ).

Figures

Fig. 10. Distribution of Parasinoennea, new genus, Sinoennea Kobelt, 1904, Pupennea, new genus, Platylennea, new genus, and Rowsonia, new genus.