Cochlostoma bicostulatum Gofas, 1989

Gofas, Serge, 2001, The systematics of Pyrenean and Cantabrian Cochlostoma (Gastropoda, Cyclophoroidea) revisited, Journal of Natural History 35 (9), pp. 1277-1369 : 1352-1353

Cochlostoma bicostulatum Gofas, 1989

Vizcaya: New material ÐAkorda: Ermita S. Pedro de Atxerre ( WP2907 ), 1 spm., UPV ; Axpe-Busturia ( WP2303 ), 3 spm., 2/3/1980, UPV ; Bilbao N: La Cadena ( WN19 ), 2 spm., UPV ; ErenÄo ( WP3102 ), 25/4/1980, UPV ; Ibarrangelua ( WP2705 ), 5 spm., 14/8/1981, UPV ; Ispazter , limestone wall, 8 spm., leg. Gofas, 15/9/1980, MNHN ; Lekeitio : Atxurras ( WP40 ), 3/1/1980, UPV ; OrdunÄa : Txarlazo ( VN 9759 (ou 94-58), 4 spm., 3/5/1981, UPV ; PenÄas de Ranero ( VN 6991, 800 m), 81 38* (2 juv.) spm., leg. Gofas and Gomez, 5/1989, MNHN ; PenÄas de Ranero ( VN 6991), 5 spm., 12/8/1981, UPV .

Santander: MNHN collectionÐ Laredo (VP70), 6 spm., leg. Bouchet 1969; BaÂdames (VP60), 4 spm., leg. Bouchet; Nacimiento de la Gandara ( VN 58), 2 spm., leg. Bouchet, 16/8/70; Nacimiento de la Gandara ( VN 58), 6 spm. (with manuscript name Cochlostoma azpeitianum Ortiz de Zarate ), coll. Staadt; Puente Viesgo: cueva el Castillo ( VN 2194), 30 spm., Dr. Obermaier, 1911.

New material ÐAlisas ( VN 4894, 500 m), 31 7* spm., leg. Gofas, 7/1991, MNHN .; Alisas ( VN 5194), 17/8/1985, UPV ; Alto de OrinÄon ( VP7206 ), 1 spm., 26/12/1985, UPV ; Bustablado ( VN 4992, 200 m), 22/7/1987, UPV ; Camijanes ( UN8098 ), 361 3* spm., leg. Gofas, 8/1990, MNHN ; Camijanes ( UN8098 ), 27/12/1987, UPV ; Castro Urdiales ( VP8204 ), 4 spm., 10/8/1981, UPV ; Comillas UP9604 ) 1 spm., 27/12/1985, UPV ; Gibaja: Ventalaperra cave ( VN 6998), 25 spm., 29/5/1983, UPV ; Islares , maritime limestone ( VP7804 ), 23 (4 juv.) spm., leg. Gofas, l0/11/1980, MNHN ; La Cabada ( VN 4399), 1 spm., leg. Gofas, 7/1991, MNHN ; La Hermida ( UN6990 ), 11* spm., leg. Gofas, 8/1990, MNHN ; La Hermida ( UN7090 ), 2 spm., 30/9/1985, UPV ; Linares, limestone ( UN 7090, 400 m), 7 spm., 20/9/1988, UPV ; OrenÄa ( VP0805 ), 1 spm., 27/11/1985, UPV ; between OrinÄon and Iseca, limestone scree ( VP7206 ), 93 (8 juv)1 83* (2 juv.) spm., leg. Gofas, 10/11/1980, and 7* spm., leg. Gofas, 5/1989, MNHN ; between Otanes and Talledo ( VN 8696), rocks in grass, 91 12* spm., leg. Gofas, 8/11/1980, MNHN ; Puente Arce ( VP 2208 ), 1 spm., 27/12/1985, UPV ; Puente Viesgo ( VN 2294), 21 10* spm., leg. Gofas, 7/1991, MNHN ; Puente Viesgo ( VN 2294), 3 spm., 28/12/1985, UPV ; Renedo ( UN9383 ), 300 m), 1 spm., 21/12/1988, UPV ; Riclones ( UN 8093, 220 m), 7 spm., 21/12/1988, UPV ; Saja: cascade ( UN9671 ), 2 spm., 29/12/1986, UPV ; Santillana del Mar ( VP1005 ), 59 (5 juv.)1 84*1 12# spm., leg. Gofas, 7/1991, MNHN ; Seldesuto , limestone ( VN 4995), the type material (115 spm.), leg. Gofas and Gomez, 5/1989, MNHN ; Seldesuto grotte 1 spm., 9/10/1987, UPV ; Soto ( VN 0167, 1100 m), 1 spm., 21 12 88, UPV ; Suances, rocks in seaside pasture ( VP1609 ), 72* spm., leg. Gofas, 11/11/1987, MNHN ; Trucios : near church ( VN 7990), 2 spm., UPV ; Valle de AsoÂn

( VN 6792), 3 spm., 16/8/1985, UPV .

Asturias: MNHN collectionÐ Des®ladero de Sella (UN38), 3 spm., leg. Bouchet, 9/1972; Puerto de Tarna (UN27), 3 spm., leg. Bouchet, 9/1972.

New material ÐArriondas ( UP 2405, 140 m), 1 spm., 23/9/1988, UPV ; Bulnes ( UN58 ), 5 spm., coll. J. Pino, Barcelona ; between Caces and Puerto, limestone ( TP0064 ), 81 24* spm., leg. Ortea and Gofas 8/1989, MNHN ; Covadonga ( UN3396 ), 11 9* spm., leg. Gofas, 7/1990, MNHN ; Covadonga , 3 km East ( UN3595 ), 61 33* spm., leg. Gofas, 7/1990, MNHN ; Covadonga , 2 km East ( UN3495 ), shaded creek, 111 43*1 8# spm., leg. Gofas, 7/1990, MNHN ; Covadonga : Elefante ( UN 3497, 260 m), 3 spm., 20/12/1988, UPV ; Covadonga : ( UN3396 , 1000 m), 4 spm., 20/12/1988, UPV ; Lago Enol ( UN3593 ), 1 spm., 2/10/1985, UPV ; El Condado ( TN 9790, 380 m), 10 spm., 17/9/1988, UPV ; El Mazuco ( UP 5005, 400 m), 1 spm., 23/9/1988, UPV ; Enol ( UN3793 , 1100 m), 31 45* spm., leg. Gofas, 7/1990, MNHN ; Enol ( UN39 , 1.300 m), 2 spm., leg. J. Gorbella, 8/1988, coll. L. Dantart, Barcelona ; Entrago ( QH 3684, 440 m), 16/9/1988, UPV ; Fries ( UP3110 ), 40 m 5 spm. 19/12/1988, UPV ; La Veguina ( UP3601 ), 2 spm., 23/9/1988, UPV ; Las Cuevas ( UN0492 , 460 m), 4 spm., 17/9/1988, UPV ; Lafranca ( UP7205 ), 10/4/1982, 20 spm., UPV ; Mere ( UP4402 ), 3 spm., 23/9/1988, UPV ; Naranjo de Bulnes ( UN5286 ), 6 spm., leg. RolaÂn 7/1983, MNHN ; Ortiguero ( UN4598 ), 71 7* spm., leg. Gofas, 8/1990, MNHN ; Panes , road to Merodio ( UN7799 ), 3 spm., leg. Gofas, 8/1990, MNHN ; Pimiango ( UP7506 , 90 m), 1 spm., 19/12/1988, UPV ; Priesca ( UN 2190, 280 m), 1 spm., 23/9/1988, UPV ; between Proaza and Trubia ( TP60 ), 5 spm., leg. RolaÂn, 7/1983, MNHN ; Puentevidosa ( UN 3086, 300 m), 3 spm., 22/9/1988, UPV ; Ribadesella ( UP3214 ), 5 spm., 25/9/1981, UPV ; Sotres , limestone ( UN 5788, 980 m), 38 spm., 20/9/1988, UPV ; Tielve ( UN 5790, 760 m), 5 spm., 9/8/1981, UPV ; Tielve ( UN 5790, 760 m), 1 spm., 20/9/1988, UPV ; Trubia ( TP60 ), 5 spm., coll. J. Pino, Barcelona ; Valdedios : Alto de la Campa, limestone ( TP9411 ), 81 12* spm., leg. Ortea and Gofas 8/1989, MNHN ; Viyao , limestone ( UP1308 ), 231 50*1 10#spm., (one sinistral), leg. Ortea and Gofas 8/1989, MNHN .

LeoÂn: MNHN collectionÐ Cain, Picos de Europa (UN48), 5 spm., leg. Bouchet, 9/1972.

New material ÐCongosto del Cares ( UN48 ), 2 spm., coll. J. Pino, Barcelona, 28/7/1986; Embalse de Porma ( UN1254 , 1020 m), 14/10/1988, UPV ; Oseja de Sajambre ( UN 3577, 840 m), 6 spm., 22/9/1988, UPV ; Santa Maria de ValdeoÂn , rocks ( UN4677 , 1120 m), 1 spm., 21/9/1988, UPV ; Verdera ( TP 9809, 350 m), 14/9/1988, UPV .

Index of localities cited in the text (* denotes samples used for allozymes)

Locality Province or deÂpartement UTM * Aisy s/ArmancËon Yonne EN97 * Alisas (pass) Santander VN 49 * AnÄisclo (canyon) Huesca BH61 * Aralar (range) Navarra WN85 * Aranzazu (sanctuary of) Guipuzcoa WN45 * Arcy s/Cure Yonne EN57 Arri (montagne d’) Haute-Garonne CH 05 * Arudy PyreÂneÂes Atlantique YN07 * Axiat ArieÁge DH03 BagneÁres Hautes-PyreÂneÂes BH66 Baigts de BeÂarn PyreÂneÂes Atlantique XP72 * Berga Barcelona DG06 Bielsa Huesca BH72 * Boitxols (pass) LeÂrida CG57 * La Cabada Santander VN 49 * Caces Asturias TP00 Camijanes Santander UN89 * Campo (canyon) Huesca BH80 * Ceresa Huesca BH71 * Collegats (canyon) LeÂrida CG38 * Covadonga Asturias UN39 * Covadonga (3 km E) Asturias UN39 * Eaux-Bonnes PyreÂneÂes Atlantiques YN16 * Enol (lake) Asturias UN39 * Formigal Huesca YN13 * Gorbea (mountain) Vizcaya WN26 * Gourette PyreÂneÂes Atlantiques YN15 Greixa Barcelona DG08 Haltza (pass) PyreÂneÂes Atlantiques XN57 * La Hermida Santander UN69 * Jou (pass) LeÂrida CG97 Lers (E Âtang de) ArieÁge CH 74 * Llobregat (sources) Barcelona DG18 * Lumbier (Foz de) Navarra XN32 * Massangis Yonne EN77 Montserrat Barcelona DG00 * OdeÂn (Sierra de) LeÂrida CG87 * Ordesa (Bujaruelo) Huesca YN32 * OrganÄa (canyon) LeÂrida CG68 * Ornolac (bridge) ArieÁge CH 94 * Ortiguero Asturias UN59 * Pau (castle) PyreÂneÂes Atlantiques YN19 * Pierre St Martin PyreÂneÂes Atlantiques XN86 PenÄagolosa CastelloÂn YK26 La Preste PyreÂneÂes-Orientales DG59 * Quillan Aude DH34 * Ranero (PenÄas de) Vizcaya VN 69 Roquefort s/Garonne Haute-Garonne CH 38 * Santillana del Mar Santander VP10 * St Martory Haute-Garonne CH 38 UrdoÂn Santander UN68 Ussat ArieÁge CH 94 * Viyao Asturias UP11