Phanaeus arletteae Arnaud, 2018

Moctezuma, Victor & Halffter, Gonzalo, 2021, Taxonomic revision of the Phanaeus endymion species group (Coleoptera: Scarabaeidae), with the descriptions of five new species, European Journal of Taxonomy 747, pp. 1-71 : 6-7

Phanaeus arletteae Arnaud, 2018

Figs 17 , 18A , 19A

Phanaeus (Notiophanaeus) arletteae Arnaud, 2018: 4 .

Phanaeus blanchardi ab. viridicollis (unavailable infrasubspecific name) – Olsoufieff 1924: 149. — Edmonds 1994: 443. — Arnaud 2002b: 98. — Edmonds & Zídek 2012: 13 (as an unavailable infrasubspecific name referable to P. pyrois ).

Phanaeus (Phanaeus) blanchardi ab. viridicollis – Martínez & Pereira 1967: 68 (as synonym of P. funereus ) (unavailable infrasubspecific name).

Phanaeus (Notiophanaeus) viridicollis – Arnaud 2002b: 98–99 (as a new combination for P. blanchardi ab. viridicollis ) (unavailable infrasubspecific name). — Edmonds & Zídek 2012: 13 (as an unavailable infrasubspecific name referable to P. pyrois ).

Phanaeus viridicollis – Krajcik 2006: 153. — Edmonds & Zídek 2012: 1, 6 (as an unavailable infrasubspecific name referable to P. pyrois ).

Non Phanaeus pyrois (error) – Moctezuma & Halffter 2017: 55 (in part). — Moctezuma et al. 2017: 114, 130 (in part).

Phanaeus (Notiophanaeus) arletteae – Chamorro et al. 2018: 97; 2019: 213 , pl. 46a.

Phanaeus arletteae – Kohlmann et al. 2018: 83, 88.

Type material

Holotype (revised from photograph from the original description)

ECUADOR • ♂, Arnaud 2018: 4, pl. 1, fig. a; Guayas, Kilometric point 18 Balzar-Quevedo Road; CPFA .

Type locality

Ecuador, Guayas, Kilometric point 18 Balzar-Quevedo Road.

Distribution

Pacific slope of the Andes, south-central Ecuador ( Fig. 17 ).

Remarks

The genital morphology of this species remains unknown. We were not able to personally revise specimens of Phanaeus arletteae . Nevertheless, the external morphology of the holotype and a female paratype were illustrated by Arnaud (2018). We consider that the diagnostic characters provided by Arnaud (2018) are adequate to separate this species from closely related taxa. The commentaries of previous authors confirmed the validity of this species ( Martínez & Pereira 1967; Edmonds 1994; Arnaud 2002b; Edmonds & Zídek 2012). Particularly, the morphology of the pronotum of the female is unique within the P. endymion species group.

Figures

Fig. 17. Distribution of the Phanaeus endymion species group in South America. Phanaeus malyi Arnaud, 2001 was reported from Colombia without accurate locality. Phanaeus arletteae Arnaud, 2018 was modified from Arnaud (2018).

Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017.

Fig. 19. Elytral integument of species within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez- Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta). H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002.K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017.