Montabunus, GOODNIGHT & GOODNIGHT, 1945

Cruz-López, Jesús A. & Francke, Oscar F., 2015, Cladistic analysis and taxonomic revision of the genus Karos Goodnight & Goodnight, 1944 (Opiliones, Laniatores, Stygnopsidae), Zoological Journal of the Linnean Society 175 (4), pp. 827-891 : 856-858

publication ID 10.1111/zoj.12299

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Montabunus Goodnight & Goodnight, 1945: 2 .

Karos: Goodnight & Goodnight, 1953: 20 (in part. generic synonymy transfer by implication).

Distribution: Central part of the state of Nuevo León, Mexico.

Type species: Montabunus foliorum Goodnight & Goodnight, 1945 , by original designation.

Included species: Monotypic.

Diagnosis: Scutum quadrangular, with posterior margin rounded and wider than mid-bulge area ( Fig. 27B View Figure 27 ). Ocularium slightly displaced towards anterior margin of prosoma, low, blunt, slightly elevated above the eyes, and unarmed ( Fig. 27D, E View Figure 27 ). Opisthosoma noticeably convex in lateral view ( Fig. 27D View Figure 27 ). Posterior margin of mesotergal area III invading and dividing area IV into two parts ( Figs 27B View Figure 27 , 61I View Figure 61 ). Stigmatic area longer in males than females ( Fig. 27C View Figure 27 ). Male genitalia: pars distalis swollen in the middle in dorsal/ventral view. Apical margin reduced, slightly convex. Ten or more pairs of lateral setae in a discontinuous row, small. Six to eight ventral microsetae along lateral margins of pars distalis, basal pairs slightly longer. Three pairs of laterobasal parastylar setae. Follis slightly longer than wide. Spiniform projections visible, small. Lateral projections of follis spiniform, very long, almost as long as follis ( Fig. 28 View Figure 28 ).

Comparisons: The male genitalia in this genus are similar to those in Chapulobunus , but with the lateral projections of the follis noticeably longer in Montabunus ( Fig. 60G View Figure 60 ). In addition, the basal-most ventral microsetae are longer than the rest, unlike in Chapulobunus where they are the same size. These two genera are easily recognizable externally: Chapulobunus are robust stygnopsids, with the scutum wider, almost rounded, and have remarkable armature on mesotergal areas II and III, instead of unarmed dorsum in Montabunus .


( FIGS 27 View Figure 27 , 28 View Figure 28 , 58 View Figure 58 , 60G View Figure 60 , 61E, I View Figure 61 , 62D View Figure 62 )

Montabunus foliorum Goodnight & Goodnight, 1945: 2 , figs 4–6 (description of male and female).

Karos foliorum: Goodnight & Goodnight, 1953: 20 (by implication); Kury & Cokendolpher, 2000: 155; Kury, 2003: 238.

Type locality: MEXICO: Nuevo León: Municipio Garza García, Chipinque (25°33′N, 100°18′W) GoogleMaps .

Material examined: MEXICO: Nuevo León: 1 ♂ and 1 ♀ [paratypes CNAN (25.ix.1943; F. Bonet)] (25°33′N, 100°18′W). Municipio Garza García, Chipinque GoogleMaps . 1 ♂ and 1 ♀ [paratypes AMNH (25.ix.1943; F. Bonet)]. Same locality . 1 ♀ [ AMNH (; without col.)]. Same locality . 1 ♂ and 2 ♀ [ AMNH (; S. Peck and J. Peck)]. Same locality .

Other material: MEXICO: Nuevo León: ♂ holotype [ AMNH (15.vii.1942; C. Bolívar, F. Bonet, D. Peláez and B. Osorio)], pine and oak woods, Chipinque, Monterrey, not examined .

Diagnosis: See generic diagnosis. Redescription: Male: Measurements: scutum length 2.3, maximum scutum width 1.8. Legs. I 1.10/0.50/0.80/ 0.90, II 1.60/0.60/1.45/1.17, III 1.30/0.47/1.15/1.22, IV 1.65/0.65/1.05/1.65. Dorsum. Scutum quadrangular, width of the prosoma similar to width of mid-bulge in scutum. Opisthosoma noticeably convex in lateral view. Dorsum uniformly and densely covered by small rounded tubercles, without armature. Mesotergal area III with posterior margin invading and dividing area IV into two parts; areas III and IV slightly fused. Ocularium a little displaced from frontal margin of prosoma, small, slightly elevated above the eyes ( Fig. 27 View Figure 27 ). Lateral clear areas projected as teardrop shapes, lateral clear areas on apices of area V displaced from the vertices, in free tergites lacking lateral projections. Venter. Densely covered by rounded tubercles, similar to the dorsum. Stigmatic area shaped like an inverted ‘T’, elongated ( Fig. 27C View Figure 27 ). Coxa IV with one small dorsomesal tooth. Pedipalps. Patella with two mesal spiniform tubercles. Legs. Similar in size and ornamentation, covered by numerous rounded tubercles, femur IV slightly curved. Tarsal count 4(2):4(2):6:6. Male genitalia. Pars distalis swollen in the middle, apical margin slightly convex in the middle, short. Twelve to 13 pairs of cylindrical lateral setae, short, forming an irregular row. Three pairs of parastylar setae on the laterobasal sides of follis. Spiniform projections visible, but hidden in the follis. Six to eight pairs of ventral microsetae along lateral margins of pars distalis, basal three to four pairs longer than the rest. Lateral projections of follis spiniform, very long ( Fig. 28 View Figure 28 ). Female: Measurements: scutum length 2.3, maximum scutum width 1.8. Very similar to male, with the following differences: stigmatic area shorter in females ( Fig. 27C View Figure 27 ). Tarsal count 4(2):4(2):6:6.

Taxonomic accounts: Goodnight & Goodnight deposit- ed some paratypes of Mexican harvestmen in the Universidad Autónoma de México ( UNAM), under the care of Dr A. Hoffmann, a notable acarologist. These paratypes were originally housed in the Acarology Laboratory, Facultad de Ciencias, UNAM. Subsequently, the Instituto de Biología, UNAM ( IBUNAM) was created, and one of its main objectives is to safeguard different biological collections. Therefore, Dr Hoffmann donated all arachnid types to the Colección Nacional de Ácaros ( CNAN) ( Hoffmann, 1993). For that reason, the original repository was considered only as UNAM ( Kury, 2003). However, the CNAN now safeguards the types donated originally by Goodnight & Goodnight and has a new database of type material codes, which are different to those cited in Kury (2003).


American Museum of Natural History


Royal British Columbia Museum - Herbarium


Universidad Nacional Autonoma de Mexico


Instituto de BiIología, Universidad Nacional Autónoma de México













Cruz-López, Jesús A. & Francke, Oscar F. 2015

Karos: Goodnight & Goodnight, 1953: 20

Goodnight CJ & Goodnight ML 1953: 20

Karos foliorum:

Kury AB 2003: 238
Kury AB & Cokendolpher JC 2000: 155
Goodnight CJ & Goodnight ML 1953: 20


Goodnight CJ & Goodnight ML 1945: 2

Montabunus foliorum

Goodnight CJ & Goodnight ML 1945: 2