Cycloryx burrailensis (Godwin-Austen, 1914)

Páll-Gergely, Barna, Gojšina, Vukašin & Aravind, Neelavar Ananthram, 2025, A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae), European Journal of Taxonomy 1029, pp. 1-158 : 36-38

Cycloryx burrailensis (Godwin-Austen, 1914)

Fig. 22A–E

Alycaeus ( Cycloryx) burrailensis Godwin-Austen, 1914: 403 , pl. 147 figs 6, 6a.

Alycaeus ( Cycloryx) burrailensis – Gude 1921: 276.

Cycloryx burrailensis – Ramakrishna et al. 2010: 70.

Pincerna burrailensis – Páll-Gergely et al. 2020: 170 .

Type material examined

Lectotype (hereby selected: Fig. 22A–E)

INDIA • Japvo Peak, Naga Hills; elev. 9890 f; Godwin-Austen leg.; acc. no. 1830, NHMUK 1903.7.1.2591.1.

Paralectotypes INDIA • 5 specs; same data as for lectotype; NHMUK 1903.7 .1.2591.2 to 1903.7.1.2591.6 • several specs in two vials, one of the vials with a single shell and a pink wool; Munipur side of the Burrail;

NHMUK 1903.7.1.2606.

Type localities

“Japvo Peak, Naga Hills, 9890 ft ”; “Munipur side of the Burrail”; Angaoluo Peak.

Differential diagnosis

Densely ribbed, similar to C. granum , but has wider spaces between R3 ribs.

Remarks

In the type collection in the same box we found the three vials (1: Japvo peak, labelled as “var. minor ”, 2591; 2: several shells with label “2606”; 3: single shell, also with label “2606” and a pink wool [which usually indicates that it is the “ type ”]). Besides the size, the difference between shells from the two localities is that R3 is much more densely ribbed in “var. minor “ (from Japvo Peak).

In the original description, the type locality was given as “Japvo Peak, Naga Hills, 9890 ft ”, and it is later added that “obtained also on the Munipur side of the Burrail, No. 2606 B.M.; and Angaoluo Peak, No. 2749 B.M.”. One understanding of Godwin-Austen’s concept is to accept only the lot from Japvo Peak as syntypes, and none of the shells from the other two sites. However, in that case specimens from Burrail would be excluded from the type series, which would be strange for a species named “ burrailensis ”. Moreover, although not mentioned in the original description, the shells from Japvo Peak were labelled as “var. minor , 2591”, suggesting, that the ones from Burrail should be considered “typical”. Lastly, the description states that there is a “distant fine costulation between sutural tube and aperture”, which unambiguously refers to the larger form from Burrail (NHMUK 1903.7.1.2606). Therefore, we accept shells belonging to all three lots as syntypes.

The single shell from Angaoluo Peak was also found in the general collection, and it is similar to the ones from Burrail in size and formation of the aperture, but the R1 ribbing is approximately half as dense.

Since the syntypes were collected at three different localities, and are somewhat different in shell traits, we hereby select a lectotype to avoid further nomenclatural confusion.