Cycloryx margarita (Theobald, 1874)

Páll-Gergely, Barna, Gojšina, Vukašin & Aravind, Neelavar Ananthram, 2025, A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae), European Journal of Taxonomy 1029, pp. 1-158 : 46

Cycloryx margarita (Theobald, 1874)

Fig. 28

Alycaeus margarita Theobald in Hanley & Theobald, 1874: 39, pl. 95 fig. 10.

Alycaeus ( Cycloryx) graphiarius Godwin-Austen, 1914: 416 , pl. 146 figs 7, 7a. Syn. nov.

Alycaeus margaritus – Sowerby 1877: pl. 1, species 5.

Alycaeus ( Alycaeus) margarita – Kobelt 1902: 347.

Alycaeus ( Cycloryx) margarita – Godwin-Austen 1914: 416.

Alycaeus margarita – Gude 1921: 211–212.

Alycaeus ( Cycloryx) graphiarius – Gude 1921: 280.

Pincerna margarita – Páll-Gergely et al. 2020: 180 .

Pincerna graphiaria – Páll-Gergely et al. 2020: 177 .

Diagnosis

This species is characterized by a globular shell, and widely-spaced, strong ribs on the entire shell surface.

Type material examined

MYANMAR • 1 syntype of A. margarita ( Fig. 28A–D); Upper Salwin ; NHMUK 1888.12 .4.892 • 1 syntype of A. margarita ; Upper Salwin ; NHMUK 1888.12 .4.893 • 3 syntypes of A. graphiarius ( Fig. 28E–I, mentioned as 1 syntype in Páll-Gergely et al. 2020); Shan States; Fedden leg.; Blanford coll.; NHMUK 1906.4 .4.21 (two of them are A. margarita , the third shell belongs to another species, see remarks) .

Type locality

“ Shan Provinces” ( A. margarita ); not specified for A. graphiarius .

Remarks

Two syntypes of A. graphiarius agreed with the original description (Godwin-Austen, 1914) by having widely-spaced ribs, but the third shell had much denser ribbing, and therefore, it clearly belongs to a different species.

The two sparsely ribbed syntypes of A. graphiarius are identical to those of A. margarita in all shell characters (size, shape, density of ribs), therefore the former is a junior synonym of the latter.