Efferia exacta, Scarbrough & Perez-Gelabert, 2009

Aubrey G. Scarbrough & Daniel E. Perez-Gelabert, 2009, Review of the West Indian species of Efferia Coquillett (Diptera: Asilidae) with 13 new species and checklist: Part II. Hispaniola, Puerto Rico, and Lesser Antilles including Tobago and Trinid, Zootaxa 1994, pp. 1-66: 18-20

publication ID

http://doi.org/ 10.5281/zenodo.1455810

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scientific name

Efferia exacta

sp. nov.

Efferia exacta  sp. nov.

Figs. 21–26View FIGURES 21–26, 89View FIGURES 89–96, Table 1

Male. Measurements, body 19.0– 22.1 mm; wing 15.0–15.0 mm. Face, thorax, and sternites brownish-yellow or yellowish-brown tomentose, vestiture mostly black. Head: Face ventrally and base of palpi narrowly white setose, 1 specimen with sparse black facial bristles and palpal setae; face at antennae 1/2–2/3 as wide as 1 eye. Occiput mostly white setose. Flagellum slightly longer than scape. Occiput dorsally black setose with 9–10 stout black postocular bristles, thinner, longer black bristles extend to or beyond mid-lateral line of head.

Thorax: Pronotum dark yellowish-brown or golden brown tomentose, bristles and setae black. Scutum with wide middle stripe and 2 large lateral spots dark brown tomentose, middle stripe divided with short diffused golden brown tomentose stripe anteriorly; tomentum otherwise light to dark yellowish-brown; vestiture mostly black, sparse white setae above wing base and narrow anterior of scutum anteriorly; anterior scutal setae as long as scape; 7–8 lateral and 4–5 prominent dorsocentral bristles. Scutellum yellowish-brown tomentose with long, usually black, sometimes mixed black and yellow, setae and 6–8, usually 6 stout, marginal bristles, setae as long as scape and pedicel combined. Pleura dark golden-brown tomentose, vestiture usually entirely black.

Wing ( Fig. 89View FIGURES 89–96): Brown, sometimes darker boarding veins; costal dilation absent; calypter fringe black. Crossvein r-m just before or at apical 1/3 of cell d. Base of cell r4 beyond cell d; R4 recurrent vein slightly shorter than diameter of cell basally.

Legs: Coxae dark yellowish-brown tomentose, mostly white setose with sparse black setae, often abundant on fore coxa. Femora black, setae and bristles mostly black, long, white setae below fore femur basally. Tibiae brownish-yellow or reddish, apices black; bristles black. Tarsi black or reddish-black, bristles black, dense dark reddish-brown setae ventrally.

Abdomen: Mostly black, segment 8 reddish. Tergites 1–5 mostly dark brown tomentose, appearing black at some light angles, narrow sides yellowish-brown or brownish-yellow except white triangles or spots in apical corners of 2–4, areas of white tomentum in corners decrease in size sequentially with largest being on 2; segments 6–7 entirely white tomentose. Tergites 1–5 mostly black setose with setae extending to lateral margins of 3–5; segment 8 reddish, setae black. Sternites 1–5 dark or pale brownish-yellow tomentose. Basal 3 tergites laterally and sternites with mostly long, erect, setae, lengths decreasing beyond with shortest setae on sternites 4–7 except apical margin of sternite 8; color of setae on sternites variable ranging from entirely white, black, or mixed.

Terminalia ( Figs. 21–24View FIGURES 21–26): Mostly black, margins slightly reddish; mostly black setose, sparse yellowish setae apically; apical and basal width not significantly different, not wider apically. Epandrium 3.1–3.3 longer than wide; narrowed apically, apex without notch, angular dorsally, narrow posteriorly. Gonostylus narrowed, asymmetrically pointed apically. Aedeagus with tubes curved downward. Gonocoxa reddish basally, black setose, setae not especially dense, shorter basally than posteriorly; anterior surface oblique, not acutely pointed.

Female. Measurements, body 21.1–22.3 mm; wing 15.9–16.9 mm. Head: Face at antennae 1/2–?. Palpi mostly white setose, apical 1/3 black. Thorax: Anterior scutal setae subequal in length to scape. Scutellar setae about as long as scape. Wing: Costal cell brownish. Crossvein r-m at apical 2/5 of cell d. R4 recurrent vein subequal or longer than base of cell r4. Calyptera white setose. Legs: Coxae entirely pale setose or with sparse black setae. Abdomen: Tergites 1–7 mostly dark brown tomentose and black setose, narrow lateral margins mostly or entirely yellowish-brown tomentose, apical corners of tergite 2–4 or 5 white tomentose, areas of white tomentum decrease in size sequentially with tergite 2 being largest. Sternites entirely or medially yellowish-brown tomentose, often slightly whitish laterally; setae long and white on basal 2–3 sternites, black and short on sternites 4–6 or 7. Terminalia ( Figs. 25–26View FIGURES 21–26): Length 5.2–5.4 mm, 9 times longer than diameter at middle. Cercus 1/3 as long as tergite 9. Spermathecae oval, slightly wider apically, broadly rounded.

Types. DOMINICAN REPUBLIC: RD-248 Entrance to Sabana / Iglesia, Santiago Prov., 372 m / 19o20.196’N 70o45.483’W / 29.iv.2004, D. Perez, B. Hierro, R. Bastardo. (d) (holotype ♂, NMNH). Paratypes: Republica Dominicana / Santo Domingo [Prov.], Distrito / National, Parque Mirador del / Norte / 2.xii.2001 / Haruo Takizawa (1 ♂, MHND); 18314 (1 ♂, IIBZ); 5807 (1 ♀, IIBZ); HAITI: Bois Caradeux [Ouest dept.] / Coll. E. M. Ducasse ~ Port-au-Prince / HAITI / 7–10.ix.1934 ~ S.W. Bromley / Collection / 1955 (2 ♂, NMNH); same data except label ~ Erax fulvibarbis Macquart  / Det. S.W. Bromley 1936 (1 ♂, NMNH); H 5 ~ HAITI / xi.1929 ~ S.W. Bromley / Collection / 1955 (1 ♀, NMNH); Port-au-Prince [Ouest dept.] / Haiti, 30–31.xii. / 1921 ~ About 300 ft alt. / F. 4609 ~ Erax fulvibarbis Marq.  / Det. C.H. Curran (1 ♀, AMNH).

Distribution. Dominican Republic: Santiago and Santo Domingo provinces; Haiti: Ouest department. Captured from sea level to about 372 meters in elevation during April, November and December.

Etymology. Latin exacta  for precise or exact, refers to the relative ease of identifying this species.

Remarks. Efferia exacta  is distinguished by the characters in the key especially, its large size (19–22 mm), black setose patterns of the body, dark brown or brownish-yellow tomentum of the thorax, white tomentose spots in the apical corners of tergites 2–4 or 6, and in the combined characters of the terminalia ( Figs. 21–26View FIGURES 21–26).


Smithsonian Institution, National Museum of Natural History


American Museum of Natural History