Wilcoxia monae Wilcox,

Pollock, Darren A. & Reichert, Lisa A., 2019, Review of the Nearctic genus Wilcoxia James (Diptera: Asilidae: Stenopogoninae) with descriptions of three new species, Zootaxa 4695 (5), pp. 401-437: 407-409

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Wilcoxia monae Wilcox


2. Wilcoxia monae Wilcox 

( Figs 3–4View FIGURES 1–4, 18View FIGURES 17–25, 27View FIGURES 26–29, 42View FIGURES 41–46, 48View FIGURES 47–58, 60View FIGURES 59–60)

Wilcoxia monae Wilcox 1972: 44  . Type locality: “ California, Mono Co., Mammoth”; Poole & Gentili 1996: 64.

Types. Holotype, male ( CAS), labeled: “Mammoth, Cal. VIII-9 ’57 // J. Wilcox Coll. // [blue label] HOLOTYPE ♂ Wilcoxia monae J. Wilcox  // California Academy of Sciences Type No. 11715 // [small blue label] ♂ // ALLOTYPE IS STORED IN THE GENERAL COLLECTION”. The holotype was not examined (images studied at: http://researcharchive.calacademy.org/research/entomology/typesDB/default.asp). The allotype was examined. 

Derivation of specific epithet. Though not mentioned specifically by Wilcox (1972) the name “ monae  ” likely refers to the type locality, in Mono County, California.

Diagnosis. This species may be diagnosed from other species in the cinerea  group by the following combination of characters: scutellum with posterior margin bare, non-pollinose; scutum without several pairs of small, semicircular non-pollinose patches; wings light to medium brown.

Description. Male ( Fig. 2View FIGURES 1–4). TL 4.6–8.0 mm; wing 4.3–6.9 mm.

Head ( Fig. 18View FIGURES 17–25) black, covered in pale grey tomentum, with face (in profile) projecting slightly beyond eye; hairs of head white; lower occipital hairs relatively short, sparse; ocellar tubercle with 4 setae (ca. length of scape + pedicel), and a few hairs, light yellow in color; mystax ( Fig. 18View FIGURES 17–25) moderately dense, consisting of long setae (ca. length of postpedicel + style) and slightly shorter hairs, confined to lower ¼ of face; setae and hairs white, bases perpendicular to plane of face; hairs above mystax relatively long (ca. length of scape + pedicel), projecting downward, with bases ca. 45° or less from plane of face; palpi and proboscis black with white hairs; antennomeres brown, pollinose, scape and pedicel thinly so; pedicel with 1 or 2 long ventral setae; scape with several relatively long ventral setae; scape and pedicel with several short hairs dorsally; ratio of antennomere lengths (scape: pedicel: postpedicel: style+spine) = 8: 7: 29: 8.

Thorax black, covered in pale grey tomentum, dorsally with light brown tint; scutum with indistinctly-defined median stripe, darker brown pollinose; acrostichal setae absent; scutum surface uniform, without distinct non-pollinose or differently colored spots; scutum with dense pile of short, white hairs (shorter than scape), similar pre- and postsuturally; most specimens with 3 notopleural setae, 1–2 supraalar setae and 2 postalar setae; stoutest thoracic setae with amber cast; scutellum pale grey pollinose, tinted very light brown in some specimens, margin (1/4 to 3/4 of scutellum length) shining, bare; 6 marginal setae, shorter than length of scutellum, in roughly same plane as scutellum, converging medially; marginal hairs absent; disc of scutellum without hairs; pleura pale grey tinted yellowish brown; katatergite with about 12 long, fine yellowish setae, tips not “crinkly” (e.g., Fig. 44View FIGURES 41–46); wings infuscated, brown; costa complete around entire wing, to base; microtrichiae conspicuous and moderately dense on distal 1/3 of wing; halter with base brown and yellow, stem and knob yellow; legs ( Figs 3–4View FIGURES 1–4) non-pollinose, largely yellow; tibiae darkened distally, tarsi slightly darker than tibiae; femora with broad black band in middle, basal 1/5 and distal 1/3 yellow; leg hairs and setae white; stoutest setae with amber cast; femora dorsally with scattered, white appressed hairs (<femur width); ventrally with longer, suberect white hairs (longest> femur width); tibiae with sparse, short (<tibia width) appressed yellowish hairs, ventrally with a few longer, more erect hairs; mesotibial spine blackishbrown; protibiae without sigmoidal spine at apex.

Abdomen ( Figs 3View FIGURES 1–4, 27View FIGURES 26–29) brownish-black, tergites non-pollinose, shining, with sparse, short white hairs; segments relatively wide, 2–5 at least as wide as long; tergites 1–6 with lateral margins pale grey pollinose, expanded slightly posteriorly on tergites 4–6; sternites pale grey pollinose, with relatively dense, moderately long white setae, not noticeably differing in length from anterior to posterior sternites.

Male genitalia ( Fig. 48View FIGURES 47–58). Apex of dorsal gonocoxite process entire, not bifid; dorsal process extending to same level as ventral gonocoxite process; medial gonocoxite processes moderately stout, not acute, slightly curved towards phallus; gonostylus relatively wide, angulate, apex not truncate; phallus without subapical denticle on lateral margins; lateral margins narrowed subapically.

Female ( Fig. 4View FIGURES 1–4). TL 6.9–7.6 mm; wing 5.0–6.0 mm; features similar to male, except for the following: pollinosity with general pale yellow cast; wing with brown infuscation only slightly lighter than in males; abdominal segments 1–6 brownish black, 7–8 yellow.

Natural history. Two specimens of W. monae  have pertinent associated data ( Wilcox 1972). One has a label “Compositae” [= Asteraceae  ], and another “ Chrysothamnus viscidiflorus var. typicus  ”; the latter presumably indi- cates that the specimen was collected on/in this species of Asteraceae  (rabbitbush). No specific collection details are known about the two specimens collected in Suisun Marsh, but the local habitat differs appreciably from that in which the specimens from the Sierra Nevada were collected ( Solano Land Trust 2017). Specimens were collected from 15.vii–1.ix.

Distribution ( Fig. 60View FIGURES 59–60). This species seemingly has a restricted distribution. Most specimens of W. monae  are known from the Sierra Nevada Mountains of California (elevation> 5000 feet). A single locality is known from coastal California (Solano Co.); two specimens were collected at “Suisun Marsh” at an elevation of 22 feet. A single specimen is known from southeastern Oregon (Harney Co.).

Non-primary type material examined. UNITED STATES OF AMERICA. California. Mono Co. Mammoth [37.634953, -118.982381], 9.viii.1957, J. Wilcox, ( CASC, 18♂, 2♀; EMFC, 6♂, 1♀; CNC, 1♀).GoogleMaps  Nevada Co. 3 mi. N. Boca [39.426754, -120.089449], 23.vii.1961, F.D. Parker, ( BME, 1♀ [UC BME P 0003967]);GoogleMaps  Prosser Dam [39.378748, -120.138340], 15.vii.1966, D.R. Miller, ( BME, 1♂ [UC BME P 0003968]).GoogleMaps  Solano Co. Rush Ranch, Suisun Marsh , 38°12.656 N, 122°01.501 W [38.21093, -122.02502], 22 ft, 17.vii.2007, K. Bayless, ( EMFC, 2♂).GoogleMaps  Trinity Co. Tri-Forest Pk. [40.9939, -122.9166], 7500’, 26.vii.1972, T. Griswold, ( OSAC, 1♀ [ OSAC _0001005825].GoogleMaps  Tulare Co. Mineral King [36.449933, -118.593090], 8.viii.1959, W.E. Simonds, ( EMFC, 1♀).GoogleMaps  Nevada. Douglas Co. Topaz Lake [38.696287, -119.545257], 17.viii.1960, A.S. Menke, ( BME, 1♂).GoogleMaps  Oregon. Harney Co. T41S R35E S22, [43.090098, -118.998101], 30.vii.1979, Lightfoot & Cobb, ( EMFC, 1♂; OSAC, 1♀ [ OSAC–0001005824]).GoogleMaps  Literature records ( Wilcox 1972). California. Nevada Co. 1 mi S Hobart Mills [39.385460, -120.183544], 1.ix.1957,GoogleMaps  Chrysothamnus viscidiflorus var. typicus  ; Truckee [39.328031, -120.183312], 17.viii.1955, Compositae. Tulare Co. 0.5 mi E Smith Meadow, Nine Mile Canyon [est. 35.89444, -118.086389], 7850 ft., 5.viii.1961GoogleMaps  GoogleMaps  .


California Academy of Sciences


Canadian National Collection of Insects, Arachnids, and Nematodes


Oregon State Arthropod Collection














Wilcoxia monae Wilcox

Pollock, Darren A. & Reichert, Lisa A. 2019

Poole, R. W. & Gentili, P. 1996: 64
Wilcox, J. 1972: 44