Stenosmylinae Krüger, 1913: 23

Martins, Caleb Califre & Price, Benjamin W., 2020, An annotated and illustrated catalogue of the Osmylidae collection (Neuroptera) at the Natural History Museum, London, Zootaxa 4883 (1), pp. 1-61 : 48-57

Subfamily Stenosmylinae Krüger, 1913: 23

Original combination: Euporismus albatrox Tillyard, 1916: 41

Current combination: Euporismus albatrox Tillyard, 1916

Type locality: [“ Head of Condamine River, near Killarney, Q. ”], Australia, Queensland (state), near Killarney (city) (28º16’23.99”S, 152º26’40.42”’E), head of Condamine River (river) .

Holotype ♂ (by original designation) with labels: ( Fig. 53 )

Condition of type: good condition, abdominal apex and genitalia slide mounted and pinned together the specimen.

Missing parts: apical flagellomeres of both antennae, medial right tarsomeres, posterior right tarsomeres, posterior left tibia, posterior left tarsomeres.

Comments: Tillyard in the original description mentioned that E.J. Dumigan—probably Edward Jarrett Dumigan (1878–1969), who was teacher and entomologist specialized in Lepidoptera ( EOAS 2020 )—captured the four specimens utilized to describe the species. He also mentioned that the type and a paratype were housed in his collection, which was donated to the NHM by Mrs Patricia “Pattie” Tillyard in 1939; and the remaining two specimens were deposited in the collections of Peter Esben-Petersen and Nathan Banks.

Original combinations: Isostenosmylus fasciatus Kimmins, 1940: 185

Current combination: Isostenosmylus fasciatus Kimmins, 1940

Type locality: [“ Peru: Callanga”], Peru, Lima (department), Cañete (province), Callanga (hill) (near Coayllo (district) (12º32’S, 76º21’W )) .

Holotype ♂ (by original designation) with labels: ( Fig. 54 )

Condition of type: good condition, abdominal apex and genitalia slide mounted and pinned together the specimen.

Missing parts: apical left flagellomeres.

Comments: The specimen was part of the large McLachlan collection purchased from the nephew of Robert McLachlan in 1938.

Original combination: Isostenosmylus fusciceps Kimmins, 1940: 181

Current combination: Isostenosmylus fusciceps Kimmins, 1940

Type locality: [“ Peru: Callanga”], Peru, Lima (department), Cañete (province), Callanga (hill) (near Coayllo (district) (12º32’S, 76º21’W )) .

Holotype ♂ (by original designation) with labels: ( Fig. 55 )

Condition of type: good condition, apex and costal region of right forewing damages.

Missing parts: apical left flagellomeres, medial right femur, medial right tibia, medial right tarsomeres.

Comments: The specimen was part of the large McLachlan collection purchased from the nephew of Robert McLachlan in 1938.

Original combination: Isostenosmylus nigrifrons Kimmins, 1940: 185

Current combination: Isostenosmylus nigrifrons Kimmins, 1940

Type locality: [“ Ecuador: Intaj”], Ecuador, Imbaruba (province), Intag (region of Andes).

Holotype ♂ (by original designation) with labels: ( Fig. 56 )

Condition of type: good condition, metathorax glued on the pin, abdominal apex and genitalia slide mounted and pinned together with the specimen.

Missing parts: flagellomeres of both antennae.

Comments: The specimen was part of the large McLachlan collection purchased from the nephew of Robert McLachlan in 1938.

Original combination: Nymphes extranea Walker, 1853: 230

Current combination: Stenolysmus extraneus ( Walker, 1853)

Type locality: [locality unknown, not stated in original publication], country unknown (probably somewhere in southeastern Australia).

Holotype ♀ (by monotypy) with labels: ( Fig. 57 )

Condition of type: good condition.

Missing parts: right flagellomeres, apical left flagellomeres, posterior left femur, posterior left tibia, posterior left tarsomeres.

Original combination: Oedosmylus latipennis Kimmins, 1940: 178

Current combination: Oedosmylus latipennis Kimmins, 1940

Type locality: [“ N.S. Wales: Mt. Kosciusko”], Australia, New South Wales (state), Kosciuzko National Park (park), Mount Kosciuszko (mountain) (36°27’21”S, 148°15’48.61”E) .

Holotype ♂ (by original designation) with labels: ( Fig. 58 )

Condition of type: good condition, abdominal apex and genitalia slide mounted and pinned together with the specimen.

Missing parts: there are no missing parts.

Comments: The specimen was part of the large McLachlan collection purchased from the nephew of Robert McLachlan in 1938.

Original combination: Oedosmylus montanus Kimmins, 1940: 179

Current combination: Oedosmylus montanus Kimmins, 1940

Type locality: [“ Queensland: National Park”], Australia, Queensland (state), National Park (park) (probably the Peak Range National Park, NP 202 (22º38’S, 147º55’E )) .

Holotype ♂ (by original designation) with labels: ( Fig. 59 )

Condition of type: good condition, abdominal apex and genitalia slide mounted and pinned together with the specimen.

Missing parts: right flagellomeres, apical left flagellomeres, anterior left femur, anterior left tibia, anterior left tarsomeres.

Comments: This specimen was part of Tillyard’s collection, which was donated to the NHM by Mrs Patricia “Pattie” Tillyard in 1939.

Original combination: Osmylus pallidus McLachlan, 1863: 113

Current combination: Oedosmylus pallidus ( McLachlan, 1863)

Type locality: [“ Hab. Australia ”], probably Australia, New South Wales (state), Sydney (district) (33°51’54”S, 151°12’34”E) or Australia, Queensland (state), Moreton Bay (region) (27°17’24”S, 153°15’34”E) .

Holotype ♀ (by implicit monotypy) with labels: ( Fig. 60 )

Condition of type: bad condition, meso-metanotum with glue, left forewing slide mounted and pinned together with the specimen, left hindwing glued on its base, abdominal apex and genitalia slide mounted and pinned together with the specimen.

Missing parts: apical left flagellomeres, medial left femur, medial left tibia, medial left tarsomeres, posterior right tarsomeres.

Comments: Despite the original label of Holotype refers only to“ Australia ”, the NHM register states: “ Australia ”, “Sydney & Moreton Bay”, “The localities are Maitland “Partinston”. Moreton Bay , Wollanggong, Para-matta . Sydney.” Maitland (33°48’54”S, 151°01’4”E), Wollanggong (34°25’59”S, 150°52”59”E) and Paramatta (33°48’54”S, 151°0’4”E) are cities of New South Gales State, which are from or close to Greater Western Sydney region, Autralia ( ABS 2011); Moreton Bay (27°17’24”S, 153°15’34”E) is a locality between the cities of Brisbane and Redland, both in Queensland State, Australia (CGQ 2020). The donor is listed as “Edward Damel”, most likely Eduard C. F. Dämel a German entomologist and insect dealer in Hamburg ( Hannemann 1967).

Original combination: Osmylus tenuis Walker, 1853: 234

Current combination: Stenosmylus tenuis ( Walker, 1853)

Type locality: [“Van Dieman’s Land”], Australia, Tasmania (state).

Lectotype ♂ (by explicit designation, as “type” or “typus”) with labels: ( Fig. 61 )

Condition of type: good condition, apex of both forewings damaged, abdominal apex and genitalia slide mount-ed and pinned together with the specimen.

Missing parts: apical right flagellomeres, left pedicel, left flagellomeres, posterior right femur, posterior right tibia, posterior right tarsomeres.

Comments: In the original description, Walker (1853) mentioned three specimens, which are in NHMUK. Kimmins (1940) designated a male as “type”, which is the lectotype ( Fig. 61 ) of the species by explicit designation as “type” (article 74.6 of ICZN); the other two specimens (NHMUK010594260; NHMUK010594314) are considered paralectotypes.

Walker also mentioned in the original description that two specimens (a,b) are from Dr. Hooker’s collection, probably he was referring to Sir Joseph Dalton Hooker (1817–1911), a British botanist and explorer, and also Charles Darwin’s closest friend. He received material from Ronald Campbell Gunn who was an outstanding naturalist resident in Van Diemen’s Land ( BGS 2020).

The third specimen (c) was from Adelaide and is from Mr. Steven’s collection, probably Tillyard was referring to Samuel Stevens (1817–1899) who was a skilled and successful collector, forming extensive and important collection of British insects, especially Coleoptera and Lepidoptera, and he also was agent of Alfred Russel Wallace in the distribution of his natural history collections ( Baker 2001).

Original combination: Stenosmylus stenopterus McLachlan, 1867: 267

Current combination: Stenosmylus stenopterus McLachlan, 1867

Type locality: [“ Australia ”], Australia, probably South Australia (state).

Lectotype ♀ (by explicit designation, as “type” or “typus”) with labels: ( Fig. 62 )

Condition of type: good condition, sternum of mesothorax with glue, abdominal apex and genitalia slide mount-ed and pinned together with the specimen.

Missing parts: flagellomeres of both antennae.

Comments: In the original description, McLachlan (1867) mentioned three specimens. Kimmins (1940) designated a female as “type”, which is the lectotype ( Fig. 62 ) of the species by explicit designation as “type” (article 74.6 of ICZN); the other two specimens (NHMUK010595123; NHMUK010594709) are considered paralectotypes. Smithers (1990) mentioned “The only published precise locality in the literature appers to be that of the designated type specimen ( Kimmins 1940) from Bakewell, South Australia.”, however Bakewell refers to the collector instead of a place. The accession register notes that the specimens were donated to the NHMUK in 1859 from the collection of Robert Bakewell (1810–1867), who lived in Australia and built up a large collection of insects ( BDBC 2020).

Original combination: Stenosmylus turneri Kimmins, 1940: 171

Current combination: Stenosmylus turneri Kimmins, 1940

Type locality: [“ S.E. Queensland, Tambourine Mts.”], Australia, Queensland (state), Tamborine Mountains (mountain) (27°55’23”S, 153°10’18”E) .

Holotype ♀ (by original designation) with labels: ( Fig. 63 )

Condition of type: good condition, abdominal apex and genitalia slide mounted and pinned together with the specimen.

Missing parts: medial left femur, medial left tibia, medial left tarsomeres, posterior left femur, posterior left tibia, posterior left tarsomeres.

Comments: The specimen was collected, and probably named after Rowland Edwards Turner (1863–1945), an entomologist who worked voluntarily for the Department of Entomology of NHMUK for more than 30 years. In the inter-war period he collected for the NHMUK in Australia and South Africa ( Benson 1946).

Figures

FIGURE 53. Holotype of Euporismus albatrox Tillyard, 1916 (NHMUK010593975, male). Labels: (1) printed on white circular paper with a red edge: “Type”; (2) handwritten on faded white paper: “Killarney, Q./ E.J. Dumigan/ 8.1.14.”; (3) handwritten on faded white paper: “Euporismus albatrox n.sp. Type. R.J.T.”; (4) printed on white paper: “Brit. Mus. 1939-45.”; (5) printed on white paper: “NHMUK010593975/ also encoded in one datamatrix barcode”.

FIGURE 54. Holotype of Isostenosmylus fasciatus Kimmins, 1940 (NHMUK010594177, male). Labels: (1) printed on white circular paper with a red edge: “Type”; (2) handwritten on faded white paper: “Callanga Peru”; (3) printed with handwritten inscriptions on faded white paper: “Isostenosmylus fasciatus Kimmins/ Type Ƌ/ det. D.E.Kimmins.”; (4) printed on blue paper: “McLachlan Coll. B.M. 1938-674.”; (5) printed on white paper: “BMNH(E)1253442”; (6) printed on white paper: “NH- MUK010594177/ also encoded in one datamatrix barcode”.

FIGURE 55. Holotype of Isostenosmylus fusciceps Kimmins, 1940 (NHMUK010594311, male). Labels: (1) printed on white circular paper with a red edge: “Type”; (2) handwritten on faded white paper: “Callanga Peru”; (3) printed with handwritten inscriptions on faded white paper: “Isostenosmylus fusciceps Kimm./ Type. ³/ det. D.E. Kimmins.”; (4) printed on blue pa- per: “McLachlan Coll. B.M. 1938-674.”; (5) printed on white paper: “BMNH(E)1253441”; (6) printed on white paper: “NH- MUK010594311/ also encoded in one datamatrix barcode”.

FIGURE 56. Holotype of Isostenosmylus nigrifrons Kimmins, 1940 (NHMUK010594304, male). Labels: (1) printed on white circular paper with a red edge: “Type”; (2) handwritten on faded white paper: “Intaj Ecuador”; (3) printed with handwritten inscriptions on faded white paper: “TYPE Isostenosmylus nigriforns Kimmins ³/ det. D.E. Kimmins.”; (4) printed on blue paper: “McLachlan Coll. B.M. 1938-674.”; (5) printed on white paper: “BMNH(E)1253433”; (6) printed on white paper: “NH- MUK010594304/ also encoded in one datamatrix barcode”.

FIGURE 57. Holotype of Nymphes extranea Walker, 1853 (NHMUK010594299, female). Labels: (1) printed on white circular paper with a green edge: “Type”; (2) handwritten on faded white paper: “Nymphes extraneus, (Type) Walker.”; (3) printed on white paper: “NHMUK010594299/ also encoded in one datamatrix barcode”.

FIGURE 58. Holotype of Oedosmylus latipennis Kimmins, 1940 (NHMUK010594162, male). Labels: (1) printed on white circular paper with a red edge: “Type”; (2) handwritten on faded white paper: “Mt. Kosciusko/ N.S. Wales/ 4500 ft. 21/1/85”; (3) printed with handwritten inscriptions on faded white paper: “Oedosmylus latipennis Kimm/ Type ³/ det. D.E.Kimmins.”; (4) printed on blue paper: “McLachlan Coll. B.M. 1938-674.”; (5) printed on white paper: “NHMUK010594162/ also encoded in one datamatrix barcode”.

FIGURE 59. Holotype of Oedosmylus montanus Kimmins, 1940 (NHMUK010594279, male). Labels: (1) printed on white circular paper with a red edge: “Type”; (2) printed with handwritten inscriptions on faded white paper: “National Pk./ Q. 3000 ft. 18-12-21”; (3) printed with handwritten inscriptions on faded white paper: “Oedosmylus montanus Kimmins/ Type. ³/ det. D.E. Kimmins.”; (4) printed on faded white paper: “Brit. Mus. 1939-45.”; (5) printed on white paper: “NHMUK010594279/ also encoded in one datamatrix barcode”.

FIGURE 60. Holotype of Osmylus pallidus McLachlan, 1863 (NHMUK010594266, female). Labels: 1) printed on white circular paper with a red edge: “Type”; (2) printed on faded white paper: “58.124 Australia.” [B.M. 1858-124]; (3) handwritten on blue paper: “O? pallidus McLach.”; (4) printed on white paper: “NHMUK010594266/ also encoded in one datamatrix barcode”.

FIGURE 61. Lectotype of Osmylus tenuis Walker, 1853 (NHMUK010594263, male). Labels: (1) printed on white circular paper with a green edge: “Type”; (2) handwritten on faded white circular paper: “V.D.L”/ handwritten on the reverse side of the same paper: “44 40” [B.M. 1844-40]; (3) printed with handwritten inscriptions on faded white paper: “Osmylus tenuis Walker./ Type ³/ det. D.E. Kimmins./ 31.10.39”; (4) printed on white paper: “NHMUK010594263/ one datamatrix Code”.

FIGURE 62. Lectotype of Stenosmylus stenopterus McLachlan, 1867 (NHMUK010594315, female). Labels: (1) printed on white circular paper with a red edge: “Type”; (2) printed on faded white paper: “59.24. S. Austral. (Bakewell.)” [B.M. 1859-24]; (3) handwritten on blue paper: “S. Stenopterus”; (4) printed with handwritten on faded white paper: “Stenosmylus stenopterus McL ♀/ det. D.E. Kimmins./ Type 1/11/39”; (5) printed on white paper: “NHMUK010594315/ also encoded in one datamatrix barcode”.

FIGURE 63. Holotype of Stenosmylus turneri Kimmins, 1940 (NHMUK010594117, female). Labels: (1) printed on white circular paper with a red edge: “Type”; (2) printed on faded white paper: “AUSTRALIA: R.E. Turner./ B.M.1935-240.”; (3) print- ed on faded white paper: “S.E. Queensland. Tambourine Mts./ 2-9.iv.1935.”; (4) printed with handwritten inscriptions on faded white paper: “Stenosmylus turneri Kimm./ Type ♀/ det. D.E. Kimmins.”; (5) printed on white paper: “NHMUK010594117/ also encoded in one datamatrix barcode”.