Keratodellitha anubis, 2021

Jouault, Corentin, Maréchal, Arthur, Condamine, Fabien L., Wang, Bo, Nel, André, Legendre, Frédéric & Perrichot, Vincent, 2022, Including fossils in phylogeny: a glimpse into the evolution of the superfamily Evanioidea (Hymenoptera: Apocrita) under tip-dating and the fossilized birth-death process, Zoological Journal of the Linnean Society 194, pp. 1396-1423 : 1405-1408

KERATODELLITHA ANUBIS JOUAULT, MARÉCHAL, WANG & PERRICHOT SP. NOV.

( FIGS 3 , 4 )

Z o o b a n k r e g i s t r a t i o n: u r n: l s i d: z o o b a n k. org:act: D217B456-4EEF-4EF7-87B9-A497EE9BCA66

Holotype: NIGP174739 , preserved in a rectangular piece of amber measuring 5 mm × 5 mm × 4 mm .

Type locality and horizon: Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar; upper Albian to lower Cenomanian, mid-Cretaceous.

Etymology: Named in reference to the Egyptian jackal god of the afterlife, Anubis, protector of graves and the deceased. The specific epithet is a noun in apposition.

Diagnosis: Scape short, about twice as long as pedicel; pretarsal claws of all legs bidentate; Rs+M slightly longer than 2Rs; third submarginal cell longer than second one; 1rs-m aligned with r-rs; 2rs-m not fully tubular; second discal cell long, about three times as long as wide; hind wing with five hamuli; metasoma with cylindrical gaster.

Description: Male. Total length as preserved 4.06 mm (excluding antennae and ovipositor); fore wing length ≥ 2.2 mm; integument largely dark brown and largely glabrous; integument without pronounced sculpturing or punctation, only finely imbricate; propodeum coarsely and strongly areolate; wings clear and hyaline, veins brown to light brown in preserved colour.

Head wider than long; clypeus and frons vertical and flat; genal width at about midlength 0.1 mm; compound eye 0.41 mm long, 0.27 mm wide; ocelli protruding, large, arranged in small triangle on top of vertex above compound eyes, separated by approximately their diameter and from posterior border of head by twice their diameter. Prominent facial horn projecting to length of 0.25 mm in profile (from anterior toruli margin to apex of horn); scape longer than wide, twice as long as pedicel, 0.2 mm long, 0.07 mm wide; pedicel more than twice as long as wide; flagellomere I the longest (~ 0.25 mm long). Clypeal base indistinct from frons, without medial longitudinal ridge. Palpal formula at least 5-3. Mandible large, square in frontal view, with straight outer and apical margins, margins meeting at rounded obtuse outer angle; inner margin with four blunt teeth and with one stout and longer, pointed apical tooth.

Mesosoma laterally compressed, longer than high, 1.35 mm long, ≥ 0.8 mm high; pronotum well developed, with well-developed concave anterior surface, apparently slightly longer than wide, lateral surfaces large and slightly depressed; mesoscutum with weak notauli not meeting, mesoscutum 0.5 mm long; mesoscutellum well separated from mesoscutum, slightly convex in profile, 0.2 mm long; mesopleuron with oblique depression and small, single row of weak areolae demarcating border with pronotal lateral surface, and even weaker row of areolae along border with metepisternum; propodeum coarsely and strongly areolate, dorsal length between metanotum and articulation with petiole equal to 0.23 mm. Legs not fully preserved except right hindlegs; long, with meso- and metacoxae closer than procoxa; hind femur medially swollen.

Fore wing with costal space apically about as broad as pterostigma; pterostigma much longer than wide, tapering gradually in width in apical half to acute apex; marginal cell narrow, with vein 1Rs slightly convex; vein M+Cu forking distally to midlength of Sc+R; 1M slightly curved, forming faint angle at junction with 1Rs, 1M subequal to 1Rs; 1Rs originating basal to pterostigma at distance subequal to pterostigmal width; 1rs-m aligned with r-rs; Rs+M straight; 2Rs much longer than 2M but slightly shorter than Rs+M; 2M exceedingly short and directed posteriorly to meet lm-cu; 3M slightly convex basally, then longitudinal; 2Rs slightly curved near Rs+M, then almost straight; r-rs originating before pterostigmal midlength, weakly sinuous, elongate, slightly shorter than 2Rs; marginal cell damaged and partial; two rs-m cross-veins present, second one distinctly weaker and thinner than surrounding veins (but not nebulous); second submarginal cell with posterior border 0.7 times shorter than posterior border of third one; third submarginal cell ~1.3 times longer than second; discal cell almost forming strongly slanted rhomboid (not distinctly pentagonal owing to exceptionally short 2M), lm-cu straight, 1.3 times longer than lCu; lCu-a slightly distal with 1M; 2m-cu conspicuous and tubular; 2cu-a present, enclosing subdiscal cell. Hind wing with venation nearly complete except lacking C; R with five distal hamuli, not meeting 2Rs apically; rs-m oblique, longer than 1Rs, about twice as short as first 1M; Cu present, virtually orthogonal to M + Cu; jugal lobe lacking.

Metasoma 2.0 mm long; first segment forming trapezoidal petiole, narrower basally, 0.44 mm long; tergum I and sternum I fused, without apparent indication of individual sclerites; gaster compressed laterally, gradually and slightly tapering in height from midlength to apical segment. Parameres directed backward, keel shaped, protruding ~ 0.2 mm. Aedegus not clearly visible.

Female unknown.

Figures

Figure 3. Keratodellitha anubis. Holotype NIGP174739 (male). A, habitus in right lateral view. B, head in right lateral view. C, head in frontal view. Scale bars: 0.5 mm.

Figure 4. Keratodellitha anubis. Holotype NIGP174739 (male). A, wing. B, line drawing of wing venation with nomenclature. Scale bars: 0.5 mm.